Une nouvelle espèce de <i>Clypeina</i> (Dasycladacée) dans le Berriasien du Salève et du Jura central

(1)

Article

Reference

Une nouvelle espèce de Clypeina (Dasycladacée) dans le Berriasien du Salève et du Jura central

CAROZZI, Albert V.

CAROZZI, Albert V. Une nouvelle espèce de Clypeina (Dasycladacée) dans le Berriasien du Salève et du Jura central. Archives des Sciences, 1956, vol. 9, no. 4, p. 471-477

DOI : 10.5169/SEALS-738988

Available at:

http://archive-ouverte.unige.ch/unige:126269

Disclaimer: layout of this document may differ from the published version.

1 / 1

(2)

2YHUEHHN ^- _YDQ ^)9RF ^L9DO OHDG ^6HF +DV$LQJORQ

3LOHW

³ ⁽ (]SHULHDOWD 5DGHPDNHU

HW - / )LVFKHU ^&$HP ^=HQOUDv:

5RFKH ^- HW ^- &RXUWRLV ^5USRVHV ,QQ bLRHLLPLH $HGLF

7HXEQHU ⁾ ⁶ ⁽

0DUUp

HW $ ⁽ 0XUQHHN ^$PRFR ^*HRUQ ^RWDQ

D ^1XRYD ^6HU

7XULDQ

^* "WRH$LL"L WRS$MV OHD - © 3$L,RSDO$ =HVF$U VRXV SUHVVH

9LHQQRW%RXUJLQ

^* HV &7LDQQJQRLV SDUDVLWHV ^LIHV ^3DQWHV ^FDWH

HHHV 0DVVRQ & 3DULV WRPH

,,

=HOOQHU

^- ^$RQDOVL ^&$HPWH

$OEHUW

&DUR]]L ² OQH ^©RDHH0H ^{HVSqFH GH} &O\SHLQD IODV HIDGDFpH- GDZ OH LHULDVWHZ GD $DIqHH HW GX FLUD FHQWUDO

3RXUVXLYDQW QRWUH pWXGH GHV 'DV\FODGDFpHV GX GRPDLQH MXUDVVLHQ QRXV FRQFHQWURQV QRV UHFKHUFKHV DFWXHOOHV GDQV OH

%HUULDVLHQ GRQW ^OHV QLYHDX[ j WH[WXUH JUXPHOHXVH HW ILQHPHQW SVHXGRRROLWKLTXH ^VH UpYqOHQW WUqV ULFKHV _HQ GpEULV G$OJXHV FDOFDLUHV VLSKRQpHV

'HV pFKDQWLOORQV UHPDUTXDEOHV QRXV RQW SHUPLV ^GH GpFULUH OD IRUPH QRXYHOOH VXLYDQWH

*HQUH &O\SHLQD

0LFKHOLQ

/GSSHPD FDWLQWLOD _{Q VS}

'DV\FODGDFpH FRPSRVpH GXQ FDQDO D[LDO FDOFLILp VXSSRUWDQW

j LQWHUYDOOHV UpJXOLHUV ^GHV YHUWLFLOOHV IHUWLOHV HQ IRUPH GpFXHOOH MXVTXj ^VL[ ^GHQWUH HX[ RQW ^pWp ^REVHUYpV HQ FRQQHFWLRQ /H FDQDO D[LDO FDOFLILp HVW LPSHUIRUp ^HQWUH ^OHV YHUWLFLOOHV ^VD SDURL YDULH GpSDLVVHXU HQWUH HW PP /H ^FDQDO D[LDO YDULH UpJXOLqUHPHQW ^GH GLDPqWUH

LO DWWHLQW

VRQ PLQLPXP ^j PP ^j PLGLVWDQFH HQWUH ^OHV YHUWLFLOOHV HW VpODUJLW ^DX PD[LPXP ^j PP ^j OD KDXWHXU ^GHV YHUWLFLOOHV

&HWWH YDULDWLRQ _UpJXOLqUH GX GLDPqWUH GX FDQDO D[LDO _UDSSHOOH

XQH VXSHUSRVLWLRQ ^GH ^F{QHV ^HPERvWpV SDU OHXU ^VRPPHW ^HOOH ^HVW

XQH ^GHV FDUDFWpULVWLTXHV ^GH OD IRUPH GpFULWH ILJ HW ILJ $ /HV YHUWLFLOOHV IHUWLOHV RQW XQH DOOXUH _JpQpUDOH GpFXHOOH j IRQG SODW GRQW OD SpULSKpULH pSDLVVLH ^VH UHOqYH IRUWHPHQW YHUV ^OH KDXW

(3)

6e$1&( '8 'e&(0%5(

)LJ

HW

j

&wLSHWLD HDOLQXD Q VS

&RXSH VXED[LDOH

&RXSHV WDQJHQWLHOOHV

&RXSH REOLTXH^J ^GDQV OD SDUWLH VXSpULHXUH GXQ YHUWLFLOOH

&RXSH WUDQVYHUVDOHT GDQVS

OD SDUWLH^S VXSpULHXUH GXQ YHUWLFLOOH

&RXSHV REOLTXHV SDUWLHOOHV

&RXSHVS

WUDQVYHUVDOHVT SS

GDQV OD SpULSKpULH ^GHV YHUWLFLOOHV 5HFRQVWLWXWLRQ^S VFKpPDWLTXH SSGXQS

VHFWHXU GH FRXSH WUDQVYHUVDOH PRQWUDQW ^OD GLVSRVLWLRQT

GDQV SOXVLHXUV SODQV ^GHV^SUDPHDX[ GXQ PrPH YHUWLFLOOH^S SODQS

LQIpULHXU^S SODQ PR\HQ SODQ VXSpULHXU

5HFRQVWLWXWLRQ^S^S VFKpPDWLTXHSS

GXQH _FRXSH D[LDOH PRQWUDQW OHV YDULD WLRQV GH OD SpULSKpULHT

GXQ YHUWLFLOOH^FRXSH^FRXS HW OD UDLVRQ GH VD VRXGXUH DYHF FHX[ TXLS

OHQFDGUHQWSS

PrPH ILJXUp TXH GDQV OD ILJXUH SUp FpGHQWH

(4)

ILJ ^j ILJ $ HW ' /HV YHUWLFLOOHV DWWHLJQHQW XQ GLDPqWUH H[WpULHXU GH ^j PP ^LOV VRQW FRPSRVpV GH j UDPHDX[ UD\RQQDQWV ^ELHQ TXH OHXU H[WUpPLWp GLVWDOH VRLW VRXYHQW EULVpH LOV RQW OD IRUPH _JpQpUDOH ^GH PDVVXHV LQIOpFKLHV YHUV^OH KDXW /HXUVHFWLRQ QRUPDOHPHQW FLUFXODLUH D XQ GLDPqWUH

YDULDQW

^GH PP ^SUqV ^GH OHXU LQVHUWLRQ _VXU OH FDQDO D[LDO j PP ^GDQV OHXU SDUWLH GLVWDOH GLODWpH /pSDLVVHXU ^GH OHXU SDURL ^HVW DQDORJXH ^j FHOOH GX FDQDO D[LDO HW YDULH ^GH j

PP FRPPH FHOOHFL ^HOOH ^VH FRPSRVH GXQ _DJUpJDW GH FDOFLWH OLEURUDGLpH EUXQ IRQFp /H UHPSOLVVDJH ^GH

OLQWpULHXU

GHV UDPHDX[ ^HVW IRUPp SDU XQH PRVDwTXH ^GH FDOFLWH ODUJHPHQW FULVWDOOLVpH _RX SDU ^{OD SkWH} ^GH IRQG ^GH OD URFKH HQFDLVVDQWH ILJ $ ^j '

/D GLVSRVLWLRQ ^GHV UDPHDX[ ^HQ ^{PDVVXH HVW} ^WUqV FDUDFWpULV WLTXH HW XQLTXH ^GDQV FHWWH IRUPH HQ FH VHQV TXH ^OHV UDPHDX[

VLQVqUHQW WRXV VXU XQ ERXUUHOHW YHUWLFLOODLUH _FUHX[ PDLV LOV QH VRQW _SDV WRXV GDQV ^OH PrPH SODQ FDU ^LOV SUpVHQWHQW ^GHV LQFOLQDL

VRQV UDGLDOHV YDULDEOHV SDU UDSSRUW ^j OD[H GX FDQDO FHQWUDO /HV FRQVpTXHQFHV GH FHWWH GLVSRVLWLRQ VRQW LPSRUWDQWHV

/HV YHUWLFLOOHV IRUPpV SDU OD ^VRXGXUH ^GHV UDPHDX[ ^DX PR\HQ GXQ ^{H[FqV GH} FDOFLWH FU\SWRFULVWDOOLQH RQW XQH SpULSKpULH LUUpJXOLqUHPHQW ^RQGXOHXVH GRQW ^OHV YDULDWLRQV VLQVFULYHQW HQWUH GHX[ SODQV WDQJHQWV GLVWDQWV ^GH PP HQYLURQ ILJ HW ILJ % &HWWH YDOHXU TXL UHSUpVHQWH OpSDLV VHXU PD[LPDOH GXQ YHUWLFLOOH j VD SpULSKpULH GLPLQXH _SHX j SHX ^HQ GLUHFWLRQ _FHQWULSqWH R HOOH QH GpSDVVH SDV PP GLDPqWUH SUR[LPDO PR\HQ GXQ _UDPHDX

/HV UDPHDX[ ^GX

IDLW

^GH OHXU LQVHUWLRQ _{VXU OD} PrPH FLUFRQ IpUHQFH YHUWLFLOODLUH RQW WHQGDQFH ^j VH GpIRUPHU UpFLSUR TXHPHQW ^GDQV OHXU SDUWLH SUR[LPDOH ^HQ GLUHFWLRQ GLVWDOH OD

YDULDELOLWp

^GDQV ^OD GLUHFWLRQ ^GHV _UDPHDX[ QH VXIILW _{SDV j} FRPSHQVHU OHXU DXJPHQWDWLRQ ^GH GLDPqWUH HW ^OHV GpIRUPD WLRQV VRQW

WRXW

^DXVVL SURQRQFpHV ^/HV VHFWLRQV QRUPDOHPHQW FLUFXODLUHV GHYLHQQHQW HOOLSWLTXHV S\ULIRUPHV ^RX ^JURVVLqUH PHQW SRO\JRQDOHV ILJ HW ^j ILJ %

%LHQ _TXH GLQFOLQDLVRQ YDULDEOH ^OHV UDPHDX[ VRQW ^VRXGpV HQWUH HX[ VXU ^XQH ^DVVH] JUDQGH GLVWDQFH UDGLDOH DX PRLQV

$UFKLYHV GHV 6FLHQFHV 9RO IDVF

(5)

6e$1&( '8 'e&(0%5(

PP &HWWH VRXGXUH TXL FRQVWLWXH XQH SRUWLRQ LPSRU WDQWH ^GHV YHUWLFLOOHV HVW DVVXUpH SDU ^XQ ^H[FqV ^GH ^FDOFLWH FU\SWRFULVWDOOLQH ^j WH[WXUH ^XQLIRUPH &HSHQGDQW SDU HQGURLWV RQ SHXW REVHUYHU ^GH SHWLWHV FDYLWpV LUUpJXOLqUHV GRQQDQW j OD _PDVVH _{XQ DVSHFW} ILQHPHQW FHOOXOHX[ _ILJ %

(Q VHFWLRQ WUDQVYHUVDOH IDLEOHPHQW REOLTXH ^OHV YHUWLFLOOHV DSSDUDLVVHQW FRPPH ^GHV FRXURQQHV pSDLVVHV GDQV OHVTXHOOHV OHV VHFWLRQV GH UDPHDX[ ^VRQW GLVSRVpHV HQ UDQJpHV JURVVLqUH PHQW FRQFHQWULTXHV SRXYDQW DWWHLQGUH ^OH ^QRPEUH ^GH ^WURLV /HV UDPHDX[ RIIUHQW ^GHV VHFWLRQV HOOLSWLTXHV IRUWHPHQW GpIRUPpHV OHXU GLVSRVLWLRQ FRQFHQWULTXH UpVXOWH ^GX

IDLW

_TXH

OD VHFWLRQ FRXSH ^HQ UpDOLWp ^GHV PDVVXHV VLWXpHV GDQV GHV

SODQV GLIIpUHQWV ^ELHQ TXDSSDUWHQDQW ^j XQ PrPH YHUWLFLOOH ILJ HW ^- ⁶ WI ILJ &

/D GLVSRVLWLRQ LUUpJXOLqUH ^GHV UDPHDX[ ^GDQV XQ YHUWLHLOOH FRQGXLW _SUHVTXH WRXMRXUV ^j ^OD VRXGXUH ORFDOH HQ SpULSKpULH GH GHX[ YHUWLFLOOHV ^VXFFHVVLIV ILJ HW (Q VHFWLRQ WDQJHQWLHOOH DSSDUDLVVHQW ^DORUV ^GHV FDYLWpV LUUpJXOLqUHV SODFpHV HQWUH ^OHV SRLQWV ^GH ^VRXGXUH ^&HV FDYLWpV HQ JpQpUDO UHPSOLHV SDU XQH PRVDwTXH ^GH FDOFLWH ELHQ FULVWDOOLVpH VRQW WRXMRXUV OLPLWpHV SDU ^GHV ^VHJPHQWV ^GH FRXUEHV FRQYH[HV YHUV OHXU LQWpULHXU _ILJ HW ILJ &

/D VRXGXUH SpULSKpULTXH ^GHV YHUWLFLOOHV FRQGXLW ^j OLVROHPHQW GXQH FDYLWp DQQXODLUH LUUpJXOLqUH SODFpH HQWUH OD SpULSKpULH HW OH FDQDO D[LDO &HWWH FDYLWp UHPSOLH SDU XQ ^DJUpJDW ^GH FDOFLWH RX SDU ^OH VpGLPHQW HQFDLVVDQW SHXW SUpVHQWHU SDUIRLV TXHOTXHV ORJHV OLPLWpHV SDU ^GHV GLDSKUDJPHV ^FDOFLILpV ILJ ILJ $

/HV UDPHDX[ VRQW ^LQVpUpV VXU OD SDUWLH pODUJLH GX FDQDO D[LDO TXL SDVVH HQ IDLW ^j ^FHW HQGURLW ^j XQ ERXUUHOHW

YHUWLFLOODLUH

FUHX[ ^&H ERXUUHOHW KDXW ^GH ^j PP ^ODUJH ^GH PP ^HVW ^SHUOp H[WpULHXUHPHQW HQ FH VHQV

TXLO

SRUWH ODPRUFH ^GHV VLOORQV UDGLDX[ VpSDUDQW ^OHV GLYHUV UDPHDX[

&KDFXQ GH FHX[FL SRVVqGH XQ SRUH ^GH FRPPXQLFDWLRQ DYHF OD FDYLWp FHQWUDOH TXL

DERXWLW

^GDQV ^OH ERXUUHOHW _FUHX[ VRQ GLD PqWUH PR\HQ ^HVW ^GH PP ILJ ILJ $ /D GLVWDQFH

(6)

)LJ

&9LSHLQD FDwLLX=D _{Q VS}

$ &RXSH VXED[LDOH PRQWUDQW WURLV YHUWLFLOOHV HQ FRQQHFWLRQ ; + &RXSH^S WDQJHQWLHOOH PRQWUDQW TXDWUH YHUWLFLOOHV ^HQ ^FRQQHFWLRQ

[

& &RXSHS

WUDQVYHUVDOHJJ

GDQV OD SDUWLHT

VXSpULHXUH GXQ YHUWWHLOOH ;[

' &RXSH WUDQVYHUVDOH GDQV OD SpULSKpULHS S

GXQ YHUWLHLOOH ;

(7)

6e$1&( '8 'e&(0%5(

HQWUH ^OHV SODQV ^PpGLDQV ^GH ^GHX[ ERXUUHOHWV VXFFHVVLIV HVW GH

j PP FHX[FL ^QH SDUDLVVHQW SDV SOXV DFFHQWXpV GDQV OHXU SDUWLH ^LQIpULHXUH ^RX VXSpULHXUH

5DSSRUWV ^HI ^F==pUH©FHV ² ^{/D IRUPH} TXH QRXV YHQRQV ^GH GpFULUH HVW GpILQLH SDU ODVVRFLDWLRQ ^GHV WURLV FDUDFWqUHV VXLYDQWV

9DULDWLRQ _UpJXOLqUH GX GLDPqWUH GX FDQDO FHQWUDO

,QFOLQDLVRQ YDULDEOH ^GHV UDPHDX[ HQ PDVVXH GDQV XQ PrPH YHUWLFLOOH

)RUWH LQIOH[LRQ YHUV ^OH KDXW HW JUDQGH pSDLVVHXU ^GH OD SpUL SKpULH ^GHV YHUWLFLOOHV HQ pFXHOOH

/D UHFRQVWLWXWLRQ VFKpPDWLTXH ILJ HW H[SOLTXH ^OH QRP ^GHVSqFH

WLUp

GX

ODWLQ

0LLQXXV VLJQLILDQW SHWLWH ^pFXHOOH

&HWWH IRUPH ^VH GLVWLQJXH DLVpPHQW ^GHV DXWUHV HVSqFHV GpFULWHV HW HQ SDUWLFXOLHU ^GH ^FHOOHV ^GX -XUDVVLTXH VXSpULHXU HW GX &UpWDFp LQIpULHXU TXL RQW WRXWHV ^GHV YHUWLFLOOHV ^IRUPpV ^GH UDPHDX[ SODFpV GDQV XQ ^PrPH SODQ ^& ©UDVVWFD )DYUH HW ^& PRSLZDID )DYUH PRQWUHQW ^HQ RXWUH ^GHV GpIRUPDWLRQV UpFLSURTXHV HQWUH UDPHDX[

FRQGXLVDQW j GHV VHFWLRQV VXEUHFWDQJXODLUHV ^/HV GLIIpUHQFHV VRQW pYLGHQWHV DYHF ^& ^=XFDV= (PEHUJHU TXL ^{D GHV} YHUWLFLOOHV ^GLVFRw GDX[ HW DYHF & <DEH HW 7R\DPD GRQW ^OHV

YHUWL

FLOOHV VRQW HQ FRUEHLOOHV WUqV SURIRQGHV ^& SDPU=D &DUR]]L SRVVqGH ^GHV YHUWLFLOOHV HQ HQWRQQRLU HW ^& ^PDUWH== (PEHUJHU PRQWUH ^GHV UDPHDX[ IDLEOHPHQW UHQIOpV HW

MRLQWLIV

^VHXOHPHQW VXU ^OH WLHUV SUR[LPDO ^GH OHXU ORQJXHXU

/D IRUPH pWXGLpH SUpVHQWH GDQV ^GHV FRXSHV REOLTXHV YRL

VLQHV GX FDQDO FHQWUDO ^GHV DQDORJLHV DYHF GHV FRXSHV G$FILZR

SRUH==D SRF=R=LFD

$OWK

^FHOD SURYLHQW ^GXQH VLPLOLWXGH ^GH OLQVHU WLRQ ^GHV _UDPHDX[ _VXU OH ERXUUHOHW

YHUWLFLOODLUH

^(Q UHYDQFKH OHV GLIIpUHQFHV VRQW LPSRUWDQWHV ^FKH]

$

^SRF=R=WFD ^OHV UDPHDX[

VRQW IXVLIRUPHV ^FDOFLILpV LQGLYLGXHOOHPHQW HW ^VRXGpV HQWUH HX[

VHXOHPHQW GDQV OHXU SDUWLH GLVWDOH LOV FRQVWLWXHQW ^GHV YHUWLFLOOHV pWRLOpV UHSRVDQW GLUHFWHPHQW ^OHV XQV VXU ^OHV DXWUHV VDQV 7LQWHU PpGLDLUH GXQ FDQDO FHQWUDO FDOFLILp

77R=RIWSH ² /H KRORW\SH GpVLJQp HVW UHSUpVHQWp SDU ^OHV ILJXUHV HW $ _{FRXSH Q} 30 FROOHFWLRQ &DUR]]L

,QVWLWXW

^GH *pRORJLH ^GH

O8QLYHUVLWp

^GH *HQqYH

(8)

$IWFUR=RUH ^DVVRFLpH ²OFL]RSZH==D SRGR=LFD

$OWK

^9XPHULD

=LDFRZLFD 'HHFNH $D=SLQJRSRUH==D DUPXDWD &DUR]]L $FLFD=DULD H=RLJDOD &DUR]]L &ULS]RSRUH==D SHURUD=LVVLPD &DUR]]L

OLQVWDQW

FHWWH IRUPH

D pWp UHQFRQWUpH ^GDQV ^OH %HUULDVLHQ ^GHV ORFDOLWpV VXLYDQWHV 6DOqYH &DUULqUH LQIpULHXUH ^GH 0RQQHWLHU H[WUrPH ^KDVH ^GX

QLYHDX ^j ^=9DLUD ^=HRLD=]DQ ^GH OD _FRXSH GH ( -RXNRZVN\

HW - )DYUH _ILJ _S

-XUD FHQWUDO 3Up0DULOOLHU ^SUqV ^GH 9DXPDUFXV GDSUqV ^GHV PDWpULDX[ LQpGLWV ^GH 0 +DDUVPD

1RXV UHPHUFLRQV OH 0XVpXP G+LVWRLUH QDWXUHOOH GH *HQqYH HQ

SDUWLFXOLHU

⁰⁰ ^- ^{)DYUH HW )} ^/DQWHUQR

GDYRLU

^PLV ^j QRWUH GLVSRVLWLRQ OHV FRXSHV PLQFHV VH UDSSRUWDQW DX 6DOqYH HW 0 0 +DDUVPD GH QRXV DYRLU DXWRULVp j UHSUHQGUH OpWXGH ^GH FHUWDLQV PDWpULDX[ ^GH VD WKqVH 1RWUH JUDWLWXGH YD pJDOHPHQW j 0 *)

(OOLRWW

^GH

O,UDT

3HWUROHXP &RPSDQ\

/WG

/RQGRQ SRXU ^VHV UHQVHLJQHPHQWV ELEOLRJUDSKLTXHV DLQVL TXj 00 ⁵ 0X UDW HW $ 3HUUHQRXG SRXU OHXU DLGH GDQV OD SUpSDUDWLRQ ^GHV PLFURSKRWRV

%,%/,2*5$3+,(

)DYUH - ©3UpVHQFH GXQH QRXYHOOH HVSqFH GDOJXH FDOFDLUH VLSKRQpH^{GDQV OH}9DODQJLQLHQ ^GX -XUD FHQWUDO &=MSHWQD LQRSLQDDª )FRJDH *HRRJLFDH HHH]LDH SS

GH *HQqHH PpP Q

0RUHOOHW

,,

^ª

)URF GDS OHDG SS SS

QV=W=D= GH *HRORJLH

=QLFHUVL=H GH *HQpHH