A sub-proteome of Arabidopsis thaliana mature stems trapped on Concanavalin A is enriched in cell wall glycoside hydrolases.
Texte intégral
(2)
(3) "!$#&%'( )*+,-./,02143657879:<; =>;@?A7B6?78CD EGF #H2EI=)*7J 5 K. L M-%NO,-A)* PRQ(S8TU VWYX[Z\UGZ ] ^S8_ U`S8aS )*,-N b,-)c ,Y%% d e+f `g K g h+<ge+
(4) +gieO &jk lm1e b In
(5) &1NOg +. . op\qIrYs/tuvs\uxw<ymz'{}|~umrA- -rb> Iz'&Aw"Y\qgpOs\uz'\"qIbsh\YqrqIr>Ouxrbs<Iz=tuxwL b|ou(rLs u-p<ArYs\uvs y ¡-¢&r>p\qIr>pOuq\£"uvp|vpOuv¤L|x|=|xrbuvqI¥&¦ s`gu§m<'/rmuvps\rb| \|vr¨wL YqIwb \"\qIpOspOhux©ªAbzO¨[p<\g \ ¬«rbu~s g¥ ¤(qIz++®(¯<°±¬²³YqrLuv|x|G ¤b+Y´Ozµ qrYs\w < °\¢«qg¶ rbwb<¤Nb|x|A|xrbuq@ª>«\usrb|~um@r>guvp\swL Y·¬|x<²hY>gr>\´OzO¸t¹¨»º\º ¼±¤2¬¨³«¥¸hs\uAYqIu§gHr>A| «r>r- uvqI¥ ½p|vp< I¾¦ ¦ ¦Izn°\¼¿¤ Lhuvs¿\À£[p\qI\À¨[pzÁ£¼\°±¡\záÂ<°±\¥ ¤rª`grYs>g¥ ½&pO|vp=Irbsz µ qIrbswb< Â<¢Ä¦gs+` uÅm\/r uvp\s\rb|\Æ|xrǨ[bwb YqIw È"\qIp\s\p\/ux©ªAbzɤbsª qI¾/r uvp\srb|É\Æ|xrǨÊbwb \YqIwb «wbuvbsªgu~¶ ux©ª z'¸³s\uAqIu§gHrYqIu˦Iz¦ s`gu§m<Hr> uvp\s\rb|&"\qIp\s\p+hux©ªA&rq6um¥ ÊqIus\p\s\z|xr¬µ8YqIhhª tp<|vp\s\zÌ¡\¡Ì\¯huv¶ ¥0 qI¥`Í">mgzªµ qIrYswb< ¼\¢"rOpOqr> p\uqIÎ+Ï {s<·bhp+|pOuxÐ\ªhwbux\¬ wb|xbux©ªAªIz2¸Ñs\uAqIu§g/&uxYq qIÎ>trYqIuxФqIuGzÌ\± £[p|ArqI¨"r ÒrbuG|vz+º(¯\¯\±HrYqIuGIz<µ qIrbs\w ¤pOqgqIª ÒpOsOuvs(/r-\ \p\qÓ u$0Ô[¸hh¦mz¦ s= u§m\H/r uvp\s\rb|&$|xr¨[ªwb YqIwb \h"\qIpOsp+hux©ªAbz'¨[p<\g$\«rYuvsªg¥g¤}(qIz+®(¯<°± ²³YqIrbuv|G|x<¤b\Y´z<µ qIrbswb< ½L|Ó<Â\ÂСÂ\¯h®(Â$Â\¯³±\ µ8rY´8Ó(Â\ÂÕ¡Â\¯/®(¬Â\¯hÌ\Ì {/rbuv|gÓ6Ö ×ØÙÚbÛÅÚÜÝbÞIß àÙ@Û§áâáÞ@àã ÛÅÚYßmÙ-ã äxß åÑæçIèIéê\ëYè ì ¥`|Owbp= \|xr>gbÒ\qIpªgYuvs=íÑYqI$ump+|vr>gbζmqIpOïîðñAò<óGôõ ö÷0ó~÷³øù\ñúxó~ñ\û\ñÃÑr>mqI/`gY Iuvs\ürL¶ ¶ uvsu§` wY (qIp+hr>gp\<qrYÒ\ <ýpOsý¤pOswbrbs\rYArb|vu~sý «YÒ+ rYqIp=Ibz»IYÒrqIr>gbþ\ÿ° Ñ¥ b|xbw> qgpOÒ\ pOqªIuG rLs ux\bsªgu~¶ ux Iuvs\ s\rbs\p ¤¥t « Gt*« rYs tÊ ³¦I¥ ½Ôµ t « ¡¯<° |Owbp\Ò\qIpªgYuvs= íhYq ux\bsªgu~¶ ux ="Ì<¼ p\¶$ <IÉÒ\qIpgu~sníÑYqIÉÒ\qIb\uxw>gb \ï=up+uvs\¶ pOqIr>guvwªnpc¹grYq >g pc \ IwYqI>gp<q`ÎÒr- ªíÑrÕrbs\R®( pO¶ \bÆíhqI$Ò\qIOuvw-gbÁ pnO$|vpwbrb|Guv· bÐuvs \/wL|x|íhrb|x|pOq[r0 \ Ò|vr<I/r/b¬qIrbs\(\Â\¯ p\¶ \ IÒ\qIpªgu~s=b|vp+s($gp/\|wp'Iux\ <O<qIp+|vrª@ g g¢¶ rb/ux|Gux<2íÑu§ Ip\np\¶N b Òp=IuA|~Éuvs<Op+|Ab¹uvsÕ n <<qp+|=Iu`³p\¶"wbb|x|íhrb|x|Òp+|A@rbwbwb \rYqIux\<-&½& \ÐIwbpOs r IpOqwb|xr<p+¶ux\bsªguv¶`ux Ò\qIpªgYuvs=wbp+¬Ò\qIumI<rªgÒ=rq`m\|HrLsRIq6uvs¬Ò\qIpªgbr Iª0=Ô \YqÒ\qIpªgYuvs=rYq Ò\qIb\uvw>g p OÉp´ux\p¥`qIbªAw>gr Iªzwbp\sªgru~sïuvsªgYqIrw> uvs<ï\p+hrbuvs=Iz¬rqI¹Òpgsª uvrb|G|vïu~s<Op+|AbÄuGs Iusrb|~|xuvs\/p\q& \rYO"rLs= s
(6) +spí³sh¶As\w>guvp\s=(½2 \umum&gp$p<q
(7) \s\p<í³|vb<N \"¶ uq- q`Yp\¶ Ò+|xrbsªHwbb|x| íÑrL|v| ì ¥`+|Owbp' |xr>gÑÒ\qgpªgYuvs=- é ç î[ð@ñAò<ó~ôõ ö÷-óv÷ øù\ñúGó~ñ\û\ñ wbb|~| íÑrb|G|vz wbp\s\wbrbsrOrb|Guvs z |~wp'Iux\ <O<qIp+|vrªbz |Owbp= \|xr>guvpOszOÒ|vrbsª gY .
(8) . èIé . . ëYè . &|rLs "`g/Ò|xrYÃ\uv¶ ¶ qIbsªqgpO|[u~sÎ\qIp<í" Ðrs\Ð\Yb|vp\Ò/bs gz' qIrYs Òp\qp\¶sª\ q uLs I"rbsÁíhr-gYq rYscwpOs`gu§m\g¹rÃÒ\ <=Iuxw rL|¬ Ò\ÒpOq h¶ pOqh \ÃÒ+|vrbs @t r Ip\qwbp+³ÒpOsLs Ip\¶`gb wbb|G|íÑrb|G|mÎrYq Òp\|A@rbwbwb \rq6ux\ª ¤rYq Òu§grrLsuAbr>A|§gz¡Ì\Ì\  \¨[pAq`IIz¡Ì\Ì<¼\¢@ª8p\|=@rbwwL rq6u\qIÒ\qIªIbs Ò$gp Ì\¯ p\¶cÒ|xrbsª wbb|v|íhrb|x| rbs\ wbp\s+ uÅm\gÇ <qb Ouv¶ ¶ qIbsª
(9) +uvs+¾p¶cÒp|Yq@ÓÉwbb|G|A|vp'bz \buGwL|x|=|vp=@ª[rbs\nÒbw- uvs=-+H|vrbsªwL|v|íÑrb|G|&Òp+|=r wbwY rYqIuv\wbp+¬Òp=Iu§guvpOsürbs\¹` q Aw0mqIwY \rbs( ªqIuvs<*Ò+|rLs \>b|vp\Ò+bsª rYs*rYqIÉ\u~¶ ¶gYqIbsª ¶mqIp<ÿp\s\¹Ò|vrbsªÎgÒbwbuG /gpÄrYspª \Y q g¤p'g\q p<z ¡Ì\Ì\ <Hp\Ò\ÒYqrs\ѵ'qz+°¯\¯\Â<¢0\¤L|v|+íÑrb|G|Ò\qIp<gYuvs+ I
(10) ¤ g¢wpOs q u<\g pÑíÑrY|v| rYqwL \uÅgbw>mqIp<qrqI uvs<p+|AbÉuvsR nqIY<A|xr> uvp\sÉp+¶"\qIp<í" Érs\É\Yb|vp+ÒhbsªgzpOq³\Y¶ bswbÐrYrbuvs=`"up uvwüpOqhr-uvp uw ` qªª $-rL| §z=°¯\¯< ¼ A-rb>>rY|mÅz=°¯\¯\±<¢-¤b|~| íhrb|v|Ñp+\uG¶`uvs\Ò\qIpªgu~s=`'wb rª|wbp=Iux\ <<qp+|vrª /g¢Iz"ª gYqIrªªIzN qIrs+ |wbp= |xrª@ªzÊ|r<ªrbsÒYqIp<´ux\rª rYqI¹uvs<p+|Abiuvs wbp\s+ q`Aw0 upOsz¹qIbhp+\b|v|Guvs\ p\qimq s\p<qÄp\¶*wbb|x|4íÑrb|~|wp+¬Òp\s\bsªI ÑYqIb\uvrÆ-ïrb|mÅzÄ¡Ì\Ì+º ¤p'g\q p<z2¡Ì\Ì\ µ q zO°¯\¯< ¼ A«< p\|~|xb¥@«+/uÅI>HrY|mÅz&¡Ì\Ì\ Ì \ÔAb|>HrL| §z\°¯\¯< ° ¨"bu§gqIzO°¯\¯\± ¢- ½& \¹bs(·L< p+¶h rbs\4 qrYs+g+|vwbp= |xrª@4 Ò=Yqg¶ rb/ux|vuG<ÎrqIRÒrYq uvw>A|Grq6|v*uGhÒp\q`grYs h¶ pOqh \ qIbp<qgrYs\uv· r>guvpOs p+¶ÃwL|x|ÁíÑrb|G|Òp+|AIr wbwb \rYqIuvªrb¶GgYqà \buq \YÒp'IuÅ uvp\ s µ'qz°¯\¯< ¼ $tuvsuxw»rLs -p<ArYs\uvs\z"°¯\¯\±<¢-½2 YĶ rb|x|¬uvsªgpÀYYqIrb|¶ rbuG|Guxªí³ \p=IÉ\um`gu~sw>guvpOs*umr<Icp\s*rb/uvspÄrbwbux I©ªALswbÿIuG/ux|xrYqIu§ uxª Ñbs<qIumr>gz ¡Ì\Ì¡ ¡Ì\Ì+® ¢-i{´p¥`\|wbrbs\r<ª r>mgrbw
(11) Ò\p\|AIr wbwb \rq6ux\ª Ò\qIp\<qI Iub|~ ¶`qIp \þspOs¥`qIbªAwbuvs\ bs\ýpOq¿`=`gu§m\gb uv\þ\qIp<Ò=Izi ª qIb|xbrªIuvs\ hp+sp'rbwbwb \rq6ux\ª0L{s\\p¥`\|w rbs\r @ªr>mgrbw
(12) $ÒpO|AIrbw wY rYqIuG\rw
(13) OOpOs<uvshrbshbs\p+¥`¶ r<I \uvps=(½2 L \rYOrÁ|xrYq uGhÒrbw-"p\sn /hp+|xbw>A|xrYqÑr@p+¶Òp\|~=@rwbwb \rYqIux\ª-8Ç \uâqIÐ\qIpÒÕpO¶ <<qIp\|vr<I< wbrYsÐqIbr
(14) ÉIphÕ +A`gu§m\gbÐs\p\s\¥gwbrYq`OpO <<qIr>g/\qpªÒ=|vuvs
(15) bngpÕíÑrY|v|&Òp+|=@rwbwb rYqIuv\<¬`Awb Ãrª ¥ rbwb>m|xz ¥ h> <| rLs ¥ ¶ Yq`A|vpO|»\qpÒ= mµ q \z»°¯\¯<¼\¢-*Ë\|pO|Awbrbs qIrYs |~wp' |Gr< <<qp+|vrª ˬ½ /g¢kwbrYs¾Y´ \uu§Ðp< ¾Ys\p+¥`\|Owbrbs\r<I*rbs\Ä qIrYs |Owbp= |Gr @crw> uu§ ux< µ q z °¯\¯<¼\¢- «+uâ+s\uv¶ uxwbrbsªÒ\qpO\qI ¬ rª¬ YsÉrbÐuvs¹Ò\qIp<gYp\huxwRrs\rb|~Aumhp¶"Ò+|xrbsª³wbb|v|írb|v|m `-rb>³>³r|mÅz °¯\¯\±<¢-¦ sªg-qIª uvs<*qIª =|§IíYqIÉp<<grYuvsbIuvs\ïw L|v|³w>A|§mqIªzw>A|§mqIÉhb\u=ýp+¶I b\|vuvs\z |xbrY<Iz> uvp|vr>gbi <\Òpwbp<m\|mIz"Ò\qIpªpOÒ\|vr `IÎrbs\iqIppªI ¤ \ur<rÉ> rL| §z"°¯\¯< ° £"p\qI\YqIuv Ð-rb|mÅz °¯\¯\  \£[pA\rYq >2rY|mÅz=°¯\¯+º =¤ \rYqIhpsªrY|mÅz=°¯\¯+º "íÑpOs >rb|mÅz'°¯\¯+º 0rL>rY|mÅz=°¯\¯\ ± \o >[rY|mÅz&°¯\¯\±<¢-8ÔÊswbb|~|&íÑrL|x|Ò\qpªgbpOÑpO¶/r-mqIn`gYírª\ @wYqIuObRuv s Yôó Yñ õÐ÷ñAøvó Yñk r>IIp\s>Hrb|mÅz+°¯\¯<¼\¢(|v|\ \<I¬`mA\uvª&íÑYqIArªIbbu§ YqpOsY|§\guvp\spO¶Hwbb|G|+íÑrb|G|Ò\qIpgu~s¶mqIp |GuAuvs\Îwbb|G|mp\qp\s-´< qIrbw- uvpOs pO¶&Ò\qpªgYuvs=¶mqIp+ Ò<qIuv¶ uxbwbb|G|=íÑrb|G|míhuÅ ÎIrb|§Ip|§\ upOs- hp<íÑ>Yqz Iuvswb¹rb|v|
(16) ¤ rYqIÉIbwqI>gbÒ\qgpªgYuvs+zN \Y wbrbsO ì ¥`|~wp' |Gr-gb4íhu§ *`rqn Awb ir< h¥ |§Awbp=IÄrbs h¥ rbs\s\p=Iïªq6uvs( \buvq¹Òrª-rY (qIp<\ bs\\p\Ò|vrªI/uxw*qI>guvw>A|A rbs\ p+|u Yqp"HrY|mÅz2¡Ì\Ì+® ¢0ª¦m& pA|v¬Òp=Iu|xNgp³ qrYÒ \b¾p\sh¤pOswbrbs\rYArb|vu~s ¤pOsh"¢íh \uxwb hu r|v w> uvs ´( qIrbw>gc¶mqIp\ ñ\û\ñ YñúGóv ñ Yû\÷0ó \õ<!ð ¬óv÷¹"r>|x4gpcuvsÄhp+|xbw>A|xªÃwbp\sªgrbuvs\uvs\ α¥ Ñ¥ rbs\s\p\ÒO(qIrbs\p= \|xz ¥ Ñ¥`|§Awbp\ÒO(qIrbs\p= \|p\qn gYqIuvwbrb|G|v¥`qL|xr>gqIªIuvªA< ¤rYqI|m-Ip\si>$rL| §z/¡Ì\Ì+® ¢0 ¨[bwbbsªg|z' \"¥`|βOwbp\Ò\qIpªgYp\hªp¶ =rbs/qIu~shrYsn ªArbs/uG|xíhYq$rbsrL|v=b Iuvs\ü¤pOsÎ «YÒ\ \rqIp'ÃrL¶ ¶ uvsu§` wb (qIpOÑr>gpO\qrYÒ\ <¶ p+|G|píÑb4\ ° h¥gL|bw> qIp\Ò\ \p<qI IuGrbsrªÎ Òbw- qIpOÑ> q` qIum`guvrbs+Ls-hrb|mÅzN°¯\¯< ¼ " *rbs(i>Ñrb|mÅzN°¯\¯\±<¢@½2 Rh r Ip\qIu§`4pO¶" \ÐÒ\qIpªgu~s=uv\bsªguv¶ uxb¹íhYq Ò\qIb\uvw>gÁ pnY´< qIrbwL|v|A|Grq"p\qhb¬qIrbs\wbp\¬Òp\s\bsªI0 ¤pOsÐÇrL¶ ¶ uvsu§`Õwb <qIphr>gp\\qIrYÒ\ <íÑr rb|mIpn Ibζ pOq \/w \rYqIrw>gYqIuGIr- uvp\süpO¶ ì ¥ |Guvs
(17) bÎ+|vAwbpOÒ\qgpªgYuvs+[p+#¶ $Yñ\ûõ<ð@ù\ñAòªôómøvó~% ÷ bú &=ñ\û÷ h r Iu >ÕrY|mÅz$°¯\¯\Â<¢krYs»pO¶ü /k¶mqIp+ rYqIuvp< ÃÒ+|xrbsªüp<qgrYs+ «\ b|v\p\s¾>ÕrL| §záÌ\Ì+® ' *ux|mIp\s¾rLs |§ rbs\s\z¡Ì\Ì+® +tuvsuGwh>}rb|mÅz°¯\¯< ¼ u8rbs\( "8I \r \z °¯\¯+º +tuvs\uxw$-rb|mÅz°¯\¯\ ± A²¬rsÁ¨[uG-r|mÅz °¯\¯\±<¢-¦ s4 \ux$íÑpOq
(18) z}íÑR \r-¹\YOL|pOÒb r¹s\>í Ò\qIpªgpOhuxw¹rÒ\Ò\qIp+rwY c grYqm u~s(*¶mqIp+ rkwYq`A\ Ò\qIpªgYuvsÃY´( qIrbw>[rYsü Iuvs\ ¤Np\sà «YÒ \rYqp'rL¶ ¶ uvsu~mÕwL (qIp+hr-gpO\qIrÒ\ <ngpÃuv+bs u~¶ p+|§|xwbb|G| íÑrL|v| ì ¥ |xu~s
(19) 4|wbp\Ò\qIpªgYuvs=-½2 \umh|~OwbpOÒ\qgpªgpO¹uGIuvs\uG¶ uxwbrbsªg| bs<qIuxwL b¶ p\qhÒ<<gr> u§AÃwbb|x| íÑrL|v|A wbp¬ÒrYqIbp$Ò<q>uvp< wbb|G|<íÑrb|G|Ò\qIpªgYp\h<-.
(20) ê<è é Gêxçê . . è . ç. ú~ñ\ûAøñAø Yð-óvñú *ux|G\¥mm<ÒAÎî[ð@ñAò<ó~ôõ ö÷-óv÷ øù\ñúGóvñ\û\ñAz' *rª-Iuv|xYí/
(21) +u @ribwbpªm(ÒbzÊíÑr<Î\qIpí³sÄuvsi \É\qIbbs\ p r> ° ¯ ¤ gp4°\° @¤ íhu§g Éri¡±4 ¥`Ò \p< p\ÒYq uvpÉr>Ρbº(¯
(22) 2{[ I-y &¦ s\¶ |vpOqªIwbbswª¹ gLhp+¶Ò+|xrbsªIÑr> r>mqI grY Y¡b® °\°wY³¢íhYqN Ib¬¶ p\qrYs\rb|~AIum- . ð xöñ+ð@ñAø~óGõ<ûõ öð0õø bóvû \øð@ñ ø÷ <ð-õ Ä÷>ø ÷õ$hY rb|~uxrbs\r tr-mqInmgLpO¶8î øù\ñúGóvñ\û\ñ¹Ñªr `qIuvs\c¡b®(¥ °\°¹wb uvsÃ|vbs<< Õr> \|vr>g/¶ |vpíÑYqIuvs\É`grY$íÑYqI. I Õ¶ p\q"rs\rb|~AIu0"Ò\Ò\qp´uvr>gb|~¡¯nÕp+¶`gb uG- AªíhYqÁ`8 Òbs\bÐu~s ¡-°R¬ïp\¶uxwbb¥ wbp|v >´( qIrbw> uvp\si<A¶`¶gYqÎrbs\*\qIu~s\bcuvs r4hp\q`grYq/í³u§ ïr¹Òª |vɶ p\qкihuvs ½ ÉY´< qIrbw- uvpOsi(=¶ ¶ q wbp\s+Ium`gRuvs °=ºÐt £[u½2qIuG[Ò ¿=ů h¤|¢Iz&°¯\¯ÐÑt ¤rb¤| zÊ¡¯ G¢"+|wYqIp+|vz&¼
(23) 2t rb¥ wbrbwp+<\|xr>gbz¡ °¯\¯ G\¢Ò\qIpªgbrª/uvs\ \uAu§gp\q"wbpw
(24) grYuv|H¥ Ì+º(Ì\Ì\z«\ur/¤ \b/uxwbrb|xz«<pumztÔz ¸ «O"¢0½& \R\qpsr>gYqIuvrb|NíÑr<wbbs q uv¶b¹míhuxwbÃr>³ ¼ -¤Æ¶ pOqÂuvs*r>ü¡¯\z~¯\¯\¯ 8zrs\4 `Òq s\r>grbs "íhrªh¶Gq \YqhwYsª qIuv¶G=4¶ p\qСbºk$uvsrYΡ<zv¯\¯\¯ &½& \ÎqIª`A|§guvs<*`ÒYqIs\r-grYsª"íÑrª I$¶ p\qwY <qIp\hr>gp\<qrYÒ\ \uxw"rYs\rb|~A@ª- õ<û[ î Göù\ñ\ð0õ<÷ñ (ó~ûó ø Yù\ð0õ ñAøvõ=ðñIöù ¡¬´k±\¥ wb wp+|§A$sÎíhr<¶`ux|G|xbníhu§g ÃÂà p+¶¤Np\sÿ«+Ò\ rYqIp=I «\ur¤N \buGwrL|xz«<N2pumz tÔz+¸«\"¢rbs íÑrªI \ íhuÅ ±Á¬ pO¶°¯Á¬t ½2qIumÒ = ¼ ѤN|â¢Iz¡¬tý¤rb¤N| xtɤ| Gts¤N| rYs/¯=ºt rª¤|+<A¶ ¶gYq-\½& \¬Ip|§+|x"Ò\qIp<gYuvs/Y´( qIrbw> Y¡¯hÑ¢&íÑrªrb\+/rbs\¬ \bs¬íhrªI Ñíhu§ ¡bºÎ¬ïp+¶ \um"<A¶`¶gYq[r>r/¶ |vp<í qIr>gpO¶ºü¬§ y +8qgpªgYuvs=NíqI/b|\gbÁíhuŠЯ= °Ðt > <|x¥ α¥ |§Awbp\ÒO(qIrbs\p=Iuv\uvs¬ \¬Irb<A¶ ¶ q-<½& \b|§'r>gNíhr wbp+|v|xbw>gb\zwbp\s\wbbs qIr0g³\ ¸h|Å qIrb¶mqIbb¥ ¤ Y¡ ¯
(25) hrY¢ «u+r¬¤ \b/uxwbrb|vz&«<p<umIzOt4Ôz\¸«"¢2gpÂ\¯\¯ rbs/\uGrb|v=@brYrbuvs=`ÊÑt qIbrzº ¬t ¤Ô z¯=m¡-°=º « «\z ¯+ű ½2q uÅgpOsˬ¥Y¡¯\¯\z¡¬t ѽµ½z=° wbrYqgq uvYqrb¬Ò \p| gª}Â\¥Y¡¯ { Ñbrb|§ \w rYqIN{qgpOÒ""¬Ñz+Ô"qIrYz<µ qIrbs\wb¢-. ú bõ<÷0ó~ôù=ô\ð0õ\ú~ñ\÷ iñ øvóbó ø~ó÷. ½& \ÉqIbrbw>guvp\sÄu´(mqIwbpOs grbu~sbc°»¬týöÑH¥`|Awbp'Iu\ª «+uv/r¤N b/uxw rL|z/«</p<uzÊtÔz ¸ O« "¢z¯=m¡Êtýrwb>gr>g<¶ ¶gYq `Ò ºÅ¯<¢Iz°Ît IpOuA rY· uG\zrbs\к(¯
(26) &*p¶Ò\qIpªgu~süY´< qIrw0uvsÎr pªgr|'Op+|§A$p¶¯=ºÑA½2 ¬qIbrbw> upOs/íÑr<wbrYq qIuvbüp(r>Â\ ¤Ä¶ p\q"º¬ pα\¯Áhuvs \YÒbs\\uvs\Ðp\s rbw> uu§ \¢rs\Á gpOÒ\ÒÊ\ "rb\\u§ up+shp\¶¯=º³Ñ ¯= ¼¬t Ip<u§A wb \|vp\qIuv<<¤p\sª qgpO|xíhqI`gp\Ò\Òb r>8 u~¬¯=¤p\s\wbbsª qr> uvp\sp\¶2 qIª =|§guvs\iö¬Ãíhr<}\>gYq huvs n Ò w> qgpOÒ\ pª p\h> qIuvwbrb|G|vÎr>¼¯+º s\ \ wbp+¬ÒArYqIumIp\sÕgpÉrÎwbrY|vuqIr>guvp\skw>q <«<grYs\rbqI¹\>uxr>guvp\s+¬OrL|Aª³¶ pOq³ÂRqYÒ+|xuxwbr>gÎrª-IrY= íÑYqI"|vª- rYs/º É !#" bú øð0õ öùõ<ðb÷-óv÷. ¦p+Y|ªw- q uvw$¶ pw>8Iuvs\ `¦ {µ ¢íÑrªÒAYqI¶ pOq b/ Iuvs\n°¼üwLÑ¥ux/hp\u~|xuv·<ÁÒ \qrbOuxbsª `¦`H"¢"`mqguÒ { hrL|Å \wbrYq[{qIp\Ò¬ hz=Ô"qrOz+µ qIrbs\wb¢2íhuv r|vu~s rYqÒ¾\qIrb\uxbsª¶mqIp\ ¼$gprYs\h°=º(¯ üp\¶Ò\qIpªgu~s/íÑYqIhrYÒ\Ò|vuxbnp\sÎrbs¦`HÈ` qguÒж p\qNuvs¥b| qIb <<qr> uvp\snuvsü t qIrbz°t <uvp<qIbrz µ ° ¤ h«zÁ¡¯*Ñt ѽ½z"° ¦`H <A¶ ¶ YqÒ ¼¥g mtbwb \uvs»> rb|mÅz"°¯\¯<¼\¢-}µp+w> Iuvs(cíÑr rbwY \uxYAb¬ Iu~s\ r[8qIpªgrYsh¦`{µÕ¤b|G| £[uvp¥¨[rb\z ÑYqIwY=|z=¤zO¸ «+"¢-<srbw>gu"qIb <<qIr>guvp\s¬íhr< ÒYq ¶ p\qIhbir-h°\ ° -¤ ªqIu~s( ¡-°* *r>º(¯² Ò\qguvpOqh pi¶ pw> Iuvs< ½p uvhÒ\qIp<rLhÒ|x¹bsª q`zN p|§grYNíÑrªuvswYqIrªIb`gYÒh\Î`gYÒ¶mqIpO º(¯gpR¡¯\zv¯<¯\¯¬² ¯=º r>8°¯\¯¬²³z+¯=º r>º(¯\¯Ñ²³z&¡ r> ¡¯\¯\¯¹²Æ \bs\z~¯\¯\¯¹²¿¶ p\qhr pªgrY|p\¶Ì<¼zv¯\¯\¯¹²Ä \¢- ¶Ggq¬¦{µ8z¦`H ` q uÒ íÑYqIR Awbwb<IuAb|~ uvsw>r>gbÎu~sú(¯n¬t ½2qIuGÒ ®® h¤|~¢z±/t qIrªz8Â\¯ +|wYqIp+|vz ° « «+z¡ h½½¶ p\q$¡bº /uvszRrbs uvs º(¯ Ñt ½&qIuGiÒ ®® ѤN|v¢IzñÈt qIrbzÃÂ\¯ +|OwbYqIp\|vzа « «zа º uvp\p+rwb>grYhux\¬¶ pOq¬¡bº/uv s % $\q Î0r|mÅz}¡Ì+®(<¢0« q uvÒ=íÑYqI¬¶Gq` \qIbrb|x pOsh p\Òp\¶ \n¡/¥ \uvw
(27) Ðbwbp\s\\uGbs+Iuvps\rb| L| `°\¼$´h°\¼Îwb¬¢íhu§ $ \b|vÒÎp\¶"¡ |píÑ¥`hb|§guvs\ÎrYrYqIp=I¬uvsЫ &« b|vbw> qpOÒ\ pOqI um¬<A¶`¶gYq `°=ºÉt ½2qIumIz¯= °Ét \|wbuGsbz¯=m¡ « «(¢-¤Np\sª uâsªAp< hL|m Y¡\¡ ½&z.
(28) ° Å ±\¿¤4b|míhuÅ hïr<wYqp'@I¥ |vu~s
(29) uvs\rYbs ¢íhYqN Ib=«YÒrYqIr> upOs¬íÑrªwbrYq qIuvbp\Hr8°¯Ñ² ¶ p\qÈ¡¾ ¿rbs\ +AIb©ª'bsª | r>*r rY´uvÑA p¶*Â\¯ h mb|vz*¡-°¯ ² p<AYq s\uv ªgzÐsª uv|R \qp+hp\Ò\ \bs\p+|<|A"¶mqIp\sª8 \rb¬qIrbwY \³ "bs\hpO¶= L| ð-õø bó~ûh÷>øñó~ûóvû µ8p+|v|vp<íhuvs\ °Ñ¥ b|xbw> qIp\Ò\ \p<qI IuGIz¹b|mÄíÑYqI `grbuvs\ íÑu§ wbp+|v|vpuv+rb|¤ppOhr<@IuGÆ|§'Æ"°=º(¯ rbwbwpOqI<uvs(ÑpÑt4bwY \uvsh>Hrb|m `°¯\¯<¼\¢- . ô YûAøvó \ó YñAøvóGõ<ûõ öð0õø bóvû\÷ò ñ\÷÷÷âö# øð0õ Lø~ð ½& \»u~s\u=uxªArb|ÎÒ\qIpgu~s gÒpIp< grbu~sb rb¶GgYqÉ°¥ b|vbw- qIpOÒ\ p<qIªIuGÉíÑYqI»Y´AwbumIb rLsÈuvs¥` | \u<`gb$í³u§ $ q <Ò IuvsÎrbwbwbp\qI\uvs</ pÎr$mgrYs\rYqI/Ò\qgpª pwYp\| «rYs gpOs\u -rL| §z=°¯\¯\Â<¢-½q`(Ò< uvwÒYÒ< u\ ¶mqIp+¿rbwb ÐÒ\qIpªgYuvsníÑYqIrbs\rb|v\· bÐ\Ãs\rbsp ¤N¥`t « xt*«Ãp\qNt³¦I¥ ½Ô[µÉt «Ãr<Ò\qIYuvp<8I| \ªwYqIuA mtu~suxw>[rb|mÅz2°¯\¯< ¼ t4bwb \uvs¹-"rb|mÅz&°¯\¯<¼\¢-=qgpªgYuvs+ÊrLsrb|=@Õ\Ãt ³¦I¥ ½ÔµÉt*« íÑYqI uv\bsªgu~¶ ux uxr r>\ p\r>gb ¤£¦ spOs qbª=s\rbsª Ò\q pªgu~s \r>gr>r< IrYqIwY \uvs\ Iuvs t h«+¤Ô½ Ò\qIp\\qIrYh ã Ú bÛxã Úbá ã ÚbÛ ã Y×IÝ ` G Ò8Ó Gííí hr> q u´ wLuxbs\w<Åwbp+ g@brYqIwb A¶ p\q6 Ib|xªwY h|~¢0ÄÔ"s\|~ hp<í/ wbp\qIª¿>´Owb b\u~s( <qª p+|v `Ò 8¯=º ¢iíÑYqI wpOsIuv+qIb¿rª*Òp'IuÅ uA qIª =|§I0 ¦I\sª uv¶ uxwbr>guvp\s¿pO¶Ò\qIpªgbuvs+iíhu§ s\rbsp ¤N¥mt*« Gt « upOs qIrYÒ\¢íhrªÒYq ¶ p\qIhbÁíÑu§ Á£[uvp<íhp
(30) = g½2 YqIhp¹IwbuxLs uv¶ uxwbz«rYsÐ-p=Iz ¸ «O"¢0½& \RÑrLuvs*IbrqIwb ÉÒrYqIrb>gYq¬íÑYqIÐh> \up+s\uvsRp<´uvr> uvp\sr<Ouv¶ ¶ YqLs urL|"Ñp+\uv¶ uxwbr>guvpOs rYsÑ q`(Ò Iu~s rªbs<·b<\Ô"shu-wb|xbrYrYíÑrrb|v|mp<íh=<½& \î Aøù\ñúxó~ñ\û\ñ Ò<qgpªgYuvs/\r-gr>ArªIíÑr \p<í¬s\|vprb\b ¶mqIp\ \ $uvÒ=þíÑY Iu§g Û à ã àGä`ã Þ ßm×mÖ`Þ à áâÙ6Ú à ¢@ ¦I\Ysª uv¶ uxwbr>guvpOs íhrª wbp\s+Iux\YqI Iu~s\uv¶ uxw rLs í¬ bsc \Ò\qIpªgYuvs+ÎíYqIiux\Ysªguv¶`ux cíhu§ r>ü|vbr<`ΰ¾<uv¶ ¶gYqIbsª q`(Ò< uw ÒYÒ< uv\<r ¶ uvq`Hwbrs\\uG\r-gbzO˳wbp<q q É¡ Å(zO° °hrYshÂ=§Â¶ pOq&qI Òbw> uOL|vhp+s\p¥ z\<uv¥&rbs³ qguv¥ wb \rYqgb ÒYÒ< uv\<rs\h\b|§gr"¤Ns 8¯=m¡ óGõ\ó~û \õ<&ð ñAø~ó Y÷ñ\û\ñú A÷ Y÷ «<\¥ wL|x|=|xrYq |p+wbrb|vuv·(r> uvp\s rbs\þ|xbs<< p\¶ Iu~s\rb|*ÒYÒ< uv\<»íÑYqI Ò\qIb\uGw-gb Iuvs< «+Ô¨[½ à×-ß ã ÚÛ <ã Ù -ã Ö ¢ÐrYsĽrYq >m ã àã Ø<ã àÞIßvÝbÛ IÞ@à ÙIß Þ ¢ G$uL|m@bs»-Îrb|mÅzn¡Ì\Ì\ {/rbs AL|m-pOs>2rL| §z°¯\¯\¯<¢-ª8qIb\uvw>guvp\s/p¶ qIrYsILh\qIrbs\p+ÑrLuvs=íhr<\pOsÊíhu§g /"qrLbhs\p+s ÙIßGÙ Þ Úb×-Ú ã b× ÙÚÛ <ã ØÚÛ ×-ÞIá§Ú<ã Þ ¢ «wY ªíÑrbw
(31) Î>rb|vz°¯\¯\Â<¢-tp\|vbw-=|xrYq¬r<<¬rbsRÒ\¦OrL|Aª íÑYqIÃwbrb|xw>=|xr>g Iuvs\É \¹r-£"uÒ\qpO\qIrb ªx Ò8Ó xí"íí Ò s\uO¥ ¬q--§¶mq +íÑrYu~ rYuv wbp+³Òp+¥ Ò ª h|v¢- hp\hp+|pOuvª¿gp pª \Yq Ò\qgpªgYuvs= íÑYqI IbrYqwY b ¶ pOq Iuvs( £h«+½ Ò\qgpO\qIrÁ ã Ú bÛGã Úá ã ÚÛ ã ×IÝ ¢ `|§IIwb =| -&rb|mÅz&¡Ì\Ì\¯<¢- ¦I\Ysª uv¶ uxwªr- uvp\shp\¶Ò\qIpªgu~s¶ rb/uG|xux2rLs ¶m=sw> upOsrL|'\p+hrbuvs=2íÑr<ÒYqI¶ pOqIhb¬Iuvs(/tÉ hu§I ªm Ò8Ó m¬\ u§@-§ux ¥6Iu8Åwb w+ux¥`uvs hp< uv¶ Iw rbs<¢ rYs/¦ sªgYq qgp «+wrYs ã Þ bÛGã Ù -ã Ø Ú Þ ß ßm× IÙÚ ¢ [A-uG|G|p+s>2rb|mÅz+°¯\¯+º ¢-\ /rbs\¤{&íYqI ¢ ¤p( us\ \pÐ-rb|mÅzN¡Ì\Ì\Ì<¢0 wb|vr<Iux¶ uxbnr wbwpOqI<uvs(ngp h¤N"oü\r>gr-ArªI ã IÙ ªã ×-ß HYqIp<´uv\rªíhqI¬s\rbbrªuvs/ \ÊHYqp´u£[rª@ ÞIßm× YÛ -Ù-àmÞ>ã Ûvà GàmÛ <ã ÛÅÚ Þ ã ¢ m£"r
(32) rL|puxw¬> rb|mÅzO°¯\¯\±<¢0 . .
(33) .
(34) . . . . . . . . . . !. -, . . . " . . . #. $#. %. & . & . '. . ". ". (. &. . +. &) ! * .
(35) ,. . 32/,. . /.. 0. !1. 04. & 5 . 67. . . . . 8:9<;>=.. 0. &
(36) 5 . . ?. &, 3@. . 1. = . 7A. & . .
(37) . ?B"C. H !. 0 ED:;*FG 3I2. '. 6 JH. -. -ð õø bó~û%Yñ\÷ \ð YûAø÷ 8q pªgu~sÎwpOswªbs qIr0 upOs/íÑrª\>gYqIhuvsb/\ \$> p+Îp\¶2£qIrb\¶ p\qI - ¡Ì\Ì\±<¢ uvs<Îp<u~s/Yq`A rb|<A/uvsh\u-p+|bhuvshY´< qIrbw- uvpOs<A¶ ¶ qrª \ grYs\\rqI= K.
(38) . . &ç vè@çê Gç@ë çç! \øð@ñ ø~óxõ<û/õ ú bõ<÷0ó~ôù=ô\ð0õ\ú~ñ\÷ Y÷<ð-õ. ï÷ø ¾øvó~÷÷ Y÷õ îøù\ñúxó~ñ\û\ñ ¦ shr"¶ uq-`Hr0xgY¬Ò<=pÁmmAªÑ ¶mqIp+ `gbÄ um pO¶(î=øù\ñúGóvñ\û\ñ$\ Iuvs\rÒ\qIpªgYp\huxw"rYÒ\Ò\qIprbwY u§Îíhr ¹sbwª-IrYq »gp gr-|~umI riÒ\qIpª pwYp\|/¶ p\qÐ \*>´( qIrbw> uvp\s¾p+¶ \ª*Ls\·bª0Ê£[r<Ib¾pOs Ò<\|GumI b/Y´\ÒAYqIuvLs grb|=\r>grp\s \"Ò<qIu~¶ uxw r> upOsp\¶ ¶mqIpÀArYqIuvp< Ò+|rLs p\q rYs @«O b|v\pOs> rb|mÅz¡Ì\Ì+® u~|mp+sÐrbsn|Å rbs\s\z¡Ì\Ì+® tuvs\uGw>rb| Åz°¯\¯< ¼ =uHrLs( "8I \rb\z°¯\¯+º ²³rbsÁ¨[uG-> rb|mÅz$°¯\¯\±<¢Iz¬íh*\>Ob|vpOÒ ri°¥6 gYÒ -´<mqrbw> uvp\s»Ò\qIpwbbªqI<h«ªgÁÐíÑYqIi\qIp<s\»uvs¾r (=¶ ¶ q wbp\sªgrYuvs\uvs\h°¯\¯$¬t ¤rb¤N| z+¶ p\|v|vp<íbÑ\$¤Np\sh¾«+bÒ\ rYqIp=I"rb¶`¶gu~su§ wb <qIp\hr>gp\\qIrYÒ\ <A(¤rb¤N| íhrª wY p'Lsürª /hp=`b¶ ¶`uxwbuxLs "Irb|§¶ p\q"¤$ Y´< qrbw0 uvp+s £[pA\rq`rY|mÅz°¯\¯+º ¢-A½2 \umÒ\qIpª pwYp\|&u` \u~¶ ¶gYqIbsª[¶mqIpO \p=I$ bRuvsÃÒ\qIYuvp<Êwbb|G|&íhrb|G|Ò\q@pªgYp\huxwÕ mA<uG `µ8bu~·Î-"rb|mÅz&°¯\¯\±<¢Ó u¢ uvs+uÅ uxrb|`gYÒÃuGÊr$\qIuvs\\u~s\ÃuvsÃr$(=¶ ¶ q"wbp\sªgrbuvs\uvs\а¯+¯Ðt ¤Nr¤N| pÐqIY|br<I/$¤ "uvs=`gbrÐpO¶}r |vp<í¾uvp+s\uxw` qYs<< ¬<A¶ ¶ Yq rb|v|~³bgp$Ò<q>bsªH¤ Ãb|\ upOs u~uv¢ YqI"uspÁ`gÒhpO¶8wbb|x|OíÑrb|G| umIp+|r> upOsngpkr>p\uvÃ|vp+p=Iuvs\k
(39) ¤ íÑbr
(40) |~O¥gOpsngp¹wbY|v|&írb|v|³ªqIuvs\Ð \wbbsª qIuv¶Gr>guvp\s4 gYÒ= qIb©ªAuqI¶ pOqwbb|v|NíÑrb|v|"up+|xr> uvp\ s uGuvuv¢¬ \Ã|Gr ``gYÒiuxrLsrb¶ ¶ uvs+uÅ` wb <qp+hr>gp\<qIrÒ\ <¹gp¹ qIrYÒ ì ¥ |Owbp= \|xr>gb Ò\qpªgYuvs=-¨[<`A|§IiI \p<í \r>Î \ïrb¶`¶ uvs\u§ wb <qpOhr-gpO\qIrÒ\ <Æ gYÒ qIª =|Ågb u~sÇr Ius\uG¶ uxwbrbs iuvs\wqIbr<I uvs ÒAbwbuv¶ uxwÈrw> uu§guxª*p¶iIYYqrL|kY´p¥ / Iuvs\¿rYq`gu~¶ uxwbuGrb|`=` qr>gª wbp\¬ÒrYqI$gp$í¬ r>&íhr hbr<`qIÁuvs$ ¬\uGrb|=@wLq A\Ò\q pªgu~s Y´< qIrbw-I½2 ªI¬uvs\wqIbr<IªOrYqIuxb ¶mqIp+ý° §¯»¶ p\q β ¥ Ñ¥´\\|p'Iux\r<I4p ±m¡É¶ pOq β¥ ÿ\|Awbp=Iuv\rª ½r>|x ¡-¢-Ô"sc \Ér<IumÃp+¶/ \uG p<=Iq`r>guvp\s\z< \uG&Ò\qIpªp+wpO|<íÑrªrb\rÒ<gbÊ prYs\rb|\·LN \ ì ¥`|~wpOÒ\qgpªgYp\"pO¶8hr>mqI gY/0 K. . \ê æ Iö#bó \ónñLø~óómø~óY÷õ ¹÷ Yð@ñú$ ú õ<÷-ó~ôÃù=ô\ð-õ\úvñ\÷b÷/ñ ø Yð õ<ûüî Göù\ñð-õ<÷Ðñ (ó~ûómø Yù\ð0õ ñAøvõ=ðñIöù |G|bs\·b\Érbw>guu§ uxªíhYq¹/r qIbóvû bó øð-õïr>$Â\ @¤zN Iu~s\ïº(¯ \ pO¶¬Ò\qpgYuvs*rbsÒ ÑH¥ |Owbp=Iuv+ rª + qIr0g - {s<·b. ¥ Ñ¥´\\|vp=Iux\r<I ¥¥ rYqIr-u~sp+¶mqIrbs\p=Iuv\r<I α ¥ Ñ¥`\|§Aw>qIp\s\uvrªI β ¥ Ñ¥`hrbs\s\p=Iuv\r<I β ¥ h¥ hrbs\sp'Iuvrª α ¥ Ñ¥`\|§Awbp=Iuv\r<I β ¥ h¥ rL|rbw>gp=Iuv\rªI α ¥ Ñ¥`rY|vrbw-gp'Iuv\r I β ¥ h¥ +|§Awp'Iuvrª α β. O« ÒwLuv¶ uxw"rbw- uuÅ ux< s\hp+| u~s hhÒ\qIpªgYuvs(¢ ¤q`A\Ò\qgpªgYuvs ¶GgYq¤p\s¬ -´< qIrbw- «YÒ\ \rYqp' Â<° ®(Ì ¡¯\ ¼\¼\° ¡¯ °\° °± ¡¯+® ¼ ¡<° ® ° ¼Ì\ ®\º ¼±+® Â\Ì\ ® °=º ® ¡. ¨[bwbp<Yq Т ¼Â ±\± Â<¼ ±<¼ º(± Ì\ ® ¼ Â<° º(Ì. ¨[r> uvp/p\¶& Òbwbuv¶ uxw rbw> uu§guxª ¤pOs³ «YÒ\ rqIp' ¤q`A\Ò\qgpªgYuvshY´< qrbw> ° ® ¼ Å ° ů ¼ ° Â=ű ±=m¡ ºº ° m¡ Â=ÅÌ.
(41) ð-õø bõ ³óñ\û\ñú ÷0ó~÷ñ ø Yð Yû\ð-óYù ûAøõ Îøù÷-õ\ú ò<úöð-õø bó~û \øð@ñ øóvû ú bõ\÷-óvôù=ô\ð0õ\ú~ñ\÷ Y÷Êò ú ø~óvû$ñ \óvûómø ù\ð-õ ñAø~õ=ðñIöù . K. &½ \"Ò\qIp<gYp\huGwrbs\rb|~AIumíÑrªÒYqI¶ pOqIhb¬Iuvs\Ò\qIpª buvs´( qIrbw>I¶mqIp\¿¡b®(¥`°\°wb r>mqI¬`gY/r> \h|vr>gh¶`|vp<íÑYqIu~s(¹`gr<+Op\³¡¯ÁÎp\¶`gb u@`'ªíÑYqI Ibζ pOq \h>´( qIrbw> upOsÎp\¶Ò\qpgYuvs- ¶GgYq¬\qIuvs\\u~s(rbs\Éwbbsª qgu~¶mr>guvp\s\z \Ðwq`A\nÒ\qgpª buvsÉY´< qIrw>wpOs grYuvs\b¹° ºk¬p\¶"Ò\qIpªgYuvsÉrª \>gYqg$uvsb¬\¬g \£qrbO¶ p\qIhh> \p Y¡Ì\Ì\±<¢- Ķ qIrbwY uvp\shp+¶= \umÒ\qIpgu~sÑ´( qIrbw> íÑrª + @bw>g³gp ¤pOsÿ«YÒ \rYqIp=Irb¶ ¶ uvs+uÅ`kwb (qIpOr>pO\qrYÒ\ <A'{|Å\gbÐÒ\qIpªgYuvs+í$YqIwbp\s\wbbs qIr>gbÃrYsR\uxrb|A@b\z qIª =|Å uvs<uvsr"¶mqrw> uvp\s/p\¶¼¯\¯ µwbp\sªgrbuvs\uvs\¬rYp<\8Â\¯\¯ µp\¶ Ò\qIpgu~sA< rb$Ò|x"wbp\sªgrbuvs\u~s\°=º(¯ ÇpO¶ÐÒ\qpgYuvs»írªi + @bw-gbÀ p ° Ñ¥ b|vbw> qpOÒ\ p<q ªIum0qpªgYuvs=RíÑYqI»\>gw>gbÀ\Çwbp+|v|vp+ux\r | µ ¤p\pOÑr@Iux |An`grbuvs\uvs\ `µ8u~ ¡-¢0 ½2 /sªAÑAq"p+¶qIªp+|b¹gÒpIíhr "rYp<\°¯\¯=+µ8uv¶G`¥IYOLs ÒpªIHqIªIp\|b$\Ñ° h¥ b|xbw> qpOÒ\ p<qIªIuGí$YqI"rbs\rb|~· Iu~s\¬t" ¬¦¥ ½Ôµ«\ÒpªIwbp\q`qI Òp\s\\u~s( pÎÒ<qgpªgYuvs= \rYuvs\Ðhp+|xbw>A|xrYqÑr@"IrL|x|xYq \rbs°¯
(42) ÑríhqI¬s\pª}rYs\rb|\· RIuvswb \YÎrYq$s\p< >´Òªw-gbn p¹wp\sªgrYuvsk ³pOsÎ \/ArªIum¬p\¶"wrL|xw>A|xr> uvp+s¬rb\¶`qpO¿bs\<"Ò\qIb\uxw>gbn pkYs\wp+\ `Awb Ò\qIpªgYuvs= tuvs\uGwÉrbs-p<ArYs\uvs\z°¯\¯\±<¢-½& \kp Yq/Ò\qgpªgYuvs=/umIu=|xkpOsÉ \ÃL|íÑYqIkrb|p rYsrL|v\·(bizN(\ªAn pcIÑrL|x|"©ªArbsªgu§ pOqhu´(mqIÕíÑu§ ipª Yq/Ò\qgpªgYuvs=h \buqnwbp\qIªhíhYqRs\p< Ius\uG¶ uxwbrbs @Nµ8u~¶Gg bs» Òp<IÐ|vpwbrb|vuv· b»r> \4OrªIuxw Iu+p+¶$ 4Ab|íÑYqIc + @bw>g pc q`(Ò< uw \u<` uvp\sÎrbs\/Ò\qIpªgYuvs=íhYqÑux\bsªguv¶`ux $ Iuvs\nsrbs\p ¤¥t*« xt*« <{rbw p+¶ L ír Y´\Ò w>gb p wYp\sªgrYuvshp\qIn \rYsp\s\ÐÒ\qIpªgYuvsiIuvswbÐÒ\qIYuvp< n mA<uv<I \p<íh r>hp=`Ѥ ¾rqInr<Iuxw `0rLÑ>>rb|mÅz=°¯\¯\±<¢-\ÄgpªgrY|s =¬AYqp\¶¡¯<°ÎOuv¶ ¶gYq@YsªÒ\qpªgYuvs+íÑrux\bsªguv¶`uxb ½r>|x¬° A½r|xª «&¡rYs\«\°r> p\s<|vuvsY¢-btrbs<p+¶= ªIÒ\qIp<gYuvs+&íÑYqIÒ\qI IYs &uvs/@YYqrL|p¶' Òp<I&qIªpO|Ob \ΰ Ñ¥ b|Gw> qIp\Ò\ \p<qIªIux \ª`guvs<ÎÒp' g¥ qIrYs+I|xr>guvp\srb|hpOuv¶ uxw r> uvp\s="`Awb ürª|~wp' |Gr- uvp\s=-AÔÊs \NAr Iup\s \<qI<`A|§IIz< \p=IgÒpIíÑYqI"wp+|v|Gw>gb$uvs gpÑ \uq`m¥ ¶ uA"\qIp<Ò=rªI \p<í³s$uvshµuvk¡ ¤pOs AYqL|vz=r<Y´Òbw>gb\z=hp=`Hp\¶ \¡bº/gÒpIr m \"r<IuxwÑIux\p+¶ Êb| wbp\sªgrbuvs\/hpOq[ rLs p\s\Ò\qgpªgYuvs= %. S IE D2. 6. 5436. pH 7. 2. 879 :. 2 3. 2<; 2=2 23 2<4 2 56 2. 5 2 4. 4 3. 3. 3 4. 4 4. 35. 5 2. 3 5. 4. 4 5. 3 3. 5. 3 3. 10. 5 4. kDa. pH 4. 7 6. 5 5. 4 5. 3. &('*)+ , ), -, . / )10 4. 1. ÚYÙ@á bàmÛvàN×-ä
(43) ßm× ÞIÛÅÚ à Û vÞ6áâÞ ßm× ×-ßmÞ@àmÛvàã Þ ÞIá Ù-àá×-Ù Þ äxßmÙ Û×-Ú× ÙÛÅÚbÞ ÙIä ÞIß ×-Ú Þ Ù6ßm×àmÞÙ6äväxÛÚbÛ ßm× Ù × -ßxÙ äGßm× à Þ Ûà`àGØbÞ@à ×äã à ÙÛÅÚbÞ Û I×-áá×-Û Ùá ×-× Ù-à àÛâÞ bá§ØÞ-ã(Û§ä ÞIÞ6Úà × à ÞIßmÞ Û Þ äm×-ß&ÚbÙIÚb× × àmÞ ÝbÞIÚ¬äm×-ß ! #" Ù6ÚÙ6á bà`Þ@à ã Ú ÞáÅÙ Þ ß IÙ0àÞ%$\ÚYØ bÞßGÛÅÚ ßmÞ äGÞIß à × ßm×Ø à×-äà × à @×-Ú Ù@Û§ÚÛ§Ú ßm× ÞIÛ§Ú ã ßGßm× àÛÚ Þ IÛß IáÞ@àßmÞ ßmÞàmÞIÚ àmÙ ÞÛ Þ Ú Û§äxÛÞ ßm× Þ6ÛÅÚ à ãh×-áÞ ØbáâÙß Ù-à à NÙIß bÞIß àÙIßmÞÛÅÚ YÛ 6Ù ßGÛ ã 'C. ;. ? . ". . . " . . ! &. . . . ;*9. @ 2 .. . ;. . ". D. D. ;. =. . 0. C. >. C. ! >. &C. . . . . @. !. ?. J . . . . . . . ". . ". . . . . ". 3. . D. 2. . *. h×>ä Þ Þ YÞIá Ù0à Ù@Ú äxÛ§ä Þ"àmÙ Þ Þ "×-Ú Þ . µ. . . 1I9. 6. ? ". . , ". . . 2. .. C. . . . 2. . = . *. 3,. =. . . . . . -. . 6 !. 3. 6. !. ". C62. ?. .
(44) óGõ\ó~û \õ<ð&ñAø~ócöð Yôó øvóGõ<û õ ÷ ò" búxú \úvñ\ðÆúxõYñúGó >ñAøvóGõ<ûþñ\û\ô ì öð-õø bó~û\÷ %. K. K. . " ú õ<÷ . úvñAø~óGõ<û õ þóvôYûAø~ó <óYô. . «Ô[¨[½zɽrYqg0m rYs\ "qIrbbhs\p+s Ò\qpO\qIr$<ÄíÑYqIÈ b p Ò\qb<uvw>i `¥ wL|x|=|xrYq |vpwbrb|Guv· r>guvp\sp+¶ ¬uv\bsªgu~¶guG/Ò\qIpªgYuvs=- «>bs `¥6Asp<\p\¶ \n¡¯<°Îuv\bsªguv¶`uxb/Ò\qIp<gYuvs+ \Т íÑYqIÒ\qIOuvw-gb$gp O¬|vp+wrL|xuv· bnuvs/ \hwbb|x|'íÑrb|G|&r> qIu´z¡Âr>& Ò+|xrªIrh/b¬qIrbs\z±uvsªgp/ \ Ys\pOÒ|vr<IhuGw*qI>guvw>A|=hz/° uvs» \ÄwY gp\Ò\|Gr I rLs  uvsÀ ÄwY \|vp\qIp\Ò+|rª`@«\u´ Ò\qpªgYuvs=RíÑYqI Ò\qIb\uvw>g pÁhbu§ \q}grq 0g$ pÁOrbw>Ap\|v<pOqp (hwbb|v|íÑrb|G| ÑpíÑ>OYqIz'Orbw>Ap+|rYqgrYq > uvs<üu s\p<íÑY|v|x¥ ª gr>|GumI b¹uvs¹Ò|xrbsªI³rbs\Õ ÁÒ\qbOuxw>guvp\s+ÑrYqIs\p<>AqÕqIb|GuxrY|G Ñrb\|Guvs\<gpOskrYs ÑYs\ w
(45) bz°¯\¯\¯<¢-&|§gp\0 YqIzr>p<<³Ì\¯ ýp\¶ \nÒ\qIpªgu~sh \rYürnÒ\qIb\uxw>gbÐ¥gYq huvsrb|ÊIuvs\r | ÒYÒ< uv\z2íh \uxwb ¹ rLs r>rb|v|p+¶ \<IÒ\qIp<gYuvs+hrbqI/grYq gbn pR ÕbwYqI-gp<q`kÒr> ªíÑrY=½íhp Ò\qIpªgYuvswbp<=|/s\p<&¬rª@Iuv+sb$gprYs<Ð +O¥ wbb|v|=|xrYqwbp\¬ÒrYq`ghbsªªA pÎ<uGwYqIYÒrbswbuxª>míÑbYs Ò\qIb\uvw>guvp\s+íÑu§ n«Ô[¨[½*rbs\ýrq >mi«>AbsÎÒ\qIp<gYuvs+íÑYqI/Ò\qIOuvw-gbÁgpÐ \rYq p<q"r$|~Owbp= \| Ò\ \p= Ò\ r- uv|uvs\p=Iu§gp+| 8¦ ¢¬rs\wY pOq-8¬¬-´\ÒAbw>gb\zrb|~|p+¶ \nuv\bs uv¶ uG ÃÒ\qIpªgYuvs+³wbp\sªgrbuvs\b ì ¥ |Owbp= \|xr>guvpOsÉIu§g<³rªÒ\qIb\uxw>gbÐ\Ð \/tÉ huÅIÒ\q@p\\qIrYh IIbë<Ò\Ò+|vbbsªgrYq k½r>|xë2¡ r> Ë"£¢- ½& \ªiqIª`A|§IÉI pí \r> \Ò\qpOÒp'bÄÒ\qIpª pwpO|rb|v|vpíÑbÄ \iumIp+|r> upOs p+¶ÎriÒ\qIpªgYuGs ¶mqIr w> uvp\sü@Ibsªguvrb|G|vÃwp+¬Òp=IbÐp\¶ ì ¥ |Guvs
(46) bn|wbp\Ò\qIpªgYuvsNgrYqg>gb/ pÃYu§ YqN /wbb|v|íhrb|x|pOq p \Ò+|vr I/r¬b¬qIrbs\< ÑgÒ+uÅgÊ \¬rYAIbs\w ¬p\¶r¬w b|~|=íÑrb|G| Ò<qIu~¶guGwr> upOsÎ`gYÒz u§ pA|v/¬s\p<gb \r- Ò\qIp\Ò\p<qm uvp\sp\¶&Ò\qIpªgYuvs=íhuÅ rÒ\qIOuvw-gbÁuvs qIrbwbb|v|=|xrYq|vpwbrb|vu~·(r> upOshíhr<OYq Î|vp<í Y¡-° ü¢ ½& \u[Ò\qIpª pwYp\| ª ÊrYÒ\ÒbrYq-rªÊrLsÕL¶ ¶ uxwbuxLs [rL|ÅgYq s\r-muA$gpÐÒ\qI>uvp< I|k\<IwYqIuAbÐÒ\qIpª pwpO|G @b ¶ p\qî øù\ñúxó~ñ\û\ñ wbY|v|&íhrb|G|Ò\qIp<gYp\huxwrLsrb|~=@ -=q>uvp< "Ò\qIp< pwYp\|GuvswL|A\Ó u¢s\p\s\¥ \ ` q`Aw> uÅA > \p+/ wb rÑrbsrL|Aump\¶¬w>A|§mqIÐb\uxr m£"p\qg\YqIuvª>³rb|mÅz°¯\¯\ }¤N rYqIhpOs ³>³rb|m&°¯\¯+º ¢Iz íÑrªI \uvs\Ðp\¶wL|x|m"w>A|§mqInuvsn|xuG©ªAuxnb\uAÈíÑu§ R@rb|§Ip\|§\guvp\s+ £[p\qI\YqIuvª>}rb|mÅz°¯\¯\  "íÑpOs >rY| O°¯\¯+º ¢}rYs\/r w>\A u~s¶ ux|§ qIr> uvp\sp¶|xbrY< £[pA\rYq 0rY| §z=°¯\¯+º &¢ uvu¢\<` q Aw0 uA¬> \p+Iz ó HwbY|v|íÑrL|x|Ò<qIuv¶ uxwbr>guvp\s\zÒ\qIuvp\q pФ Y´< qrbw0 upOsíhu§ rYqIuvp< <A¶ ¶ q ¤N \uAr<Irh>}rb|m+°¯\¯<° £[rYYq[>rY|mÅz°¯\¯\ ± =µ8buv·-rb|mÅz°¯\¯\±<¢A½2 /wY p+uxwb/p\¶r$Òrq` uvw>A|xrYqÒ\qIpªp+wp\|íhuv|G|\ÒLsÐp\sÁ rbuv¾p+¶= \`mA<ÑrYs\hp\s \Ò|vrbsª8p\q rYsÑpO¶8u~s gYqIª I ôYûAøvó \óYñAøvóGõ<ûhñ\û\ô <ûLø~óGõ<û\ñú b ú~ñ\÷@÷-ó \óYñAøvóGõ<ûõ8öð-õø ó~û\÷ K. ¦I\Ys uv¶ uxwªr- uvpOsÇp+¶ÃÒ\qIpªgYuvs ¶ rLuG|xuG rbs\ȶGAsw> uvp\srb|Î\p+rbu~s=kíhYqÄÒYqI¶ pOq Ñb Iu~s\ @YqIrb| uvp+uvs\¶ p\qI/r- uvwÉÒ\qIp\\qIrbh/ -N8qIp<gYuvs+íYqwb|xr<IuG¶guG»rbwbwbp\qI\u~s(cgpi \ÉÒ\qIb\uvw>gbĶGAsw> uvp\srb| wb|vr<[p\¶$¤ Ò\qIp\Òp'bÁ\ü-rb>>r|m `°¯\¯\±<¢ g½r>|°z&Â<¢-+8qgpªgYuvs+Ab|vp\s<uvs\ngp¹I>OLs ¶m=sw> upOsrL|wb|xr@I<íqIÁ¶ p<s\krbwbwbp\qI\uvs<Ðgpn \/Ò\q@ªIbswbnp\¶"¶GAs\w>guvp\s\rb|\p+hrbuvs="Ò\qIOuxw>gbÃrª \ªwYqIuA/u~s¬ Êtr>gq6urL|'rLs/> \p+Ó u¢Ò\qIpªgYuvs+}rbw> uvs< pOshÒp+|=@rwbwb \rYqIux\ªuvswb|A\ rLs ª gYqIrªª u~uv¢ p´ux\p¥`qIbªAw>gr IªÑrLuvs|~*uvswb|A+ÃÒ=qIp´u\rª@ªÎrbs\i¬A|§ uxwbp\Ò\ÒYq p´uv\r I< uGuvuv¢ Ò\qIpªgYuvsÊíÑu§ Éuvs gYqIrbw> uvs(\pOrbuvs=¬uvswb|A\ÎÒ\qIpªgYuvs="íhu§g ¹|xbw>guvs¹p\q"&¨N¨ |v>'wbuvsÁqIuxwb RqYÒbr> ¢ \p\hrbu~s=³rª"íÑb|G|rÊLs\·b/Îu~s\ u=uÅgpOq! u\¢"Ò\qpgYuvs¬uvs<p+|AbÃuvsIuv+srb|Guvs\ÃÒ\qIp+wªuvswb|A\ ¶grªIwbuxwb|Guvs rqIrYu~spOrb|xrbw>grsÆÒ\qIpªgYuvs=g&¢ \¢iÒ\qpªgbrªI & uv¢iÒ\qIpªgYuvs*p\¶4O>És
(47) s\p<í¬s ¶m=sw> upOs `uvuv¢i/umwbb|G|xrbsbp 0 Ñp<íh>OYqIzO \umwb|xr-Iuv¶ uxwbr- uvp\s/umÒ\qIpumIuvps\rb|&Iu~swÊ \Êup+|vp\+uxwbrb| qIp\|v¬p+¶rbs<Îp+¶ Ò\qIp<gYuvs+qIb/rbuvs=gp/¬\>gq`u~s ½r- p=Ip<Î>rb|mÅz¡Ì\Ì\<¢-<µ8p\q>´rb¬Ò|xbz rYs Ls(·b/Îp+¶ \Ð Â4¶grbhuG|~ `ËÍÂ<¢hI \p<í/Ñrb/uvs\pÉr wbuvÉ \p+hp|vpO\ÐíhuÅ \¥ h¥m´\|vp'Iux\r<I( ÑpíÑ>qIz=u§&íhr uG\bs u~¶guGr}rbsbs<·bÊ \r-Y¶`¶guGwbuxbsªg|Î <<qIp\|\·(brYqIr-u~sp |=qI<IuxªAª¶mqIp rYqIr-u~srbs=z' \ª` uvs<¬ r>8u§ íhp< q
(48) =2rªrYs ¥¥ rqIr>uGsp+¶GqIrYsp'Iux\rª@ mtu~s\uxw">HrY| §z\°¯\¯\±<¢- . .
Documents relatifs
To test whether the vesicular pool of Atat1 promotes the acetyl- ation of -tubulin in MTs, we isolated subcellular fractions from newborn mouse cortices and then assessed
Néanmoins, la dualité des acides (Lewis et Bronsted) est un système dispendieux, dont le recyclage est une opération complexe et par conséquent difficilement applicable à
Cette mutation familiale du gène MME est une substitution d’une base guanine par une base adenine sur le chromosome 3q25.2, ce qui induit un remplacement d’un acide aminé cystéine
En ouvrant cette page avec Netscape composer, vous verrez que le cadre prévu pour accueillir le panoramique a une taille déterminée, choisie par les concepteurs des hyperpaysages
Chaque séance durera deux heures, mais dans la seconde, seule la première heure sera consacrée à l'expérimentation décrite ici ; durant la seconde, les élèves travailleront sur
A time-varying respiratory elastance model is developed with a negative elastic component (E demand ), to describe the driving pressure generated during a patient initiated
The aim of this study was to assess, in three experimental fields representative of the various topoclimatological zones of Luxembourg, the impact of timing of fungicide
Attention to a relation ontology [...] refocuses security discourses to better reflect and appreciate three forms of interconnection that are not sufficiently attended to