Modélisation et Ingénierie de la Photosynthèse

HAL Id: hal-00852282

https://hal.inria.fr/hal-00852282

Submitted on 22 Aug 2013

HAL is a multi-disciplinary open access

archive for the deposit and dissemination of sci-entific research documents, whether they are pub-lished or not. The documents may come from teaching and research institutions in France or abroad, or from public or private research centers.

L’archive ouverte pluridisciplinaire HAL, est destinée au dépôt et à la diffusion de documents scientifiques de niveau recherche, publiés ou non, émanant des établissements d’enseignement et de recherche français ou étrangers, des laboratoires publics ou privés.

Modélisation et Ingénierie de la Photosynthèse

Jean-François Cornet, Olivier Bernard

To cite this version:

Jean-François Cornet, Olivier Bernard. Modélisation et Ingénierie de la Photosynthèse. Mosseri, Rémy and Jeandel, Catherine. L’énergie à découvert, 1, CNRS Eds, pp.160-161, 2013, 978-2-271-07678-6 9782271076786. �hal-00852282�

12345678962ABC9B6AD4A6CE6CB3CB58BF2927A97CB

1 1 2345678459ABC1DEF1 8A3CC381351538B31315B313C18AC1 1 DD715CB14C4 1!"F6DF#!$#7"%1&&'(1 1 5C3)* 31CB35BB313C1D+34,1-1$./1A 031/1 (214335313C14454BC1&56721%!$F1D33,1 86845ABC/A853935C/81 1 1 E B3B381$F%F:1 :B8338131F3138131 $EDEF1 1 F%#F%#DF#!"D1 $;51 '&;'(1A<1B41%5B<A BC133,1 A B3B38/*385489B58B4/81 1

441 )A BC4BA51 5)8B31 41 <38)BC1 =4) BA8381 31 5A)*83,1 <8AC1 *BA315A AB3C1 >1 AC1 3C15B334,1?A53<BA51A<B)BC4BA51A))4531C<383BCBA5@/1D=3C14 3)351 314C1<A81 41  831 3C1 )B8A4 3C1 AA1 3C1 3A8C1 B535C3C1 CA51 48A8=1B1 < A0C1 <A81 33 A<<381 3C1 )AB 3C131B338C15B334,131A)< 3,B1351A5BA513C1A*83BC1313C1BC<ACBBC131<8ABA51 A5CB8C1?<1AA*BA8438C1A1843C40C@183 334513<35451AC131 =B55B38B3/1

4A8C=B 1 C=4B1 31 <8AB831 31 41 *BA)4CC31 31 <8A1 3C1 A5B1 31 49A51 >1 31 31 AC1 3C1  )35C1 58BBC1 CAB351 )4B535C1 >1 3C1 A53584BA5C1 A<B)4 3C1 <A81 41 8ABCC453/1 43C1 3,1 <8B5B<4,1 438C1 83C45C1 ?)B53))351 348B4* 3C1 45C1 31 3)<C@1 4BCC451 C81 41 8ABCC4531 CA51 4 A8C1 41 )BB83131 413)<84831A51 3C13A BA5C18C 3511 ,1=538B31CA 4B83/1

4=3A BA51 31 41 3)<84831 <31 D831 )A BC31 <481 3C1 *B 45C1 138)B3C1 B5 451 31 840A553)35131<35451A83)35131 =34<A84BA51 B31>13C11AB,1315A AB3C131 831351 C0CB)3C1 A338C1A1 38)C/1 :3C1)AB 3C1 <38)3351 35CB31 31 <83AB81 =331 31 41 3)<84831 C81 41 3B3CC31 31 8ABCC4531 31 C81 384B53C1 84BA5C1 3 3C1 31 41 C051BC31 31 <B)35C/1 441 <8B5B<4 31 4848BCB31 3C1 )AB 3C1 31  83C1 <1AAC051B3C1 83C31 4 A8C1 31 845C381 31 )BB83141C3B5113A )3184BA553 /1D314C<313C13584 1<A8141)AB5C13,184BCA5C1E11?B@1F3C1 41 <8B5B<4 31 B)B4BA51<10CB311<8A1?A))31A1<8A1CA 4B831F4B 38C@1B1B,31A51<A81 5313)<84831A5531 3C1<38A8)453C1B5B3C131538B3C13C1<1AA*BA8438C1G1?BB@1 31 14)<1 31 84B4BA51 4<<484H1 A))31 41 <8B5B<4 31 A584B531 C81 31 )4*A BC)31 3C1 )B8A6

A845BC)3C1 ?1A8C1 A89431 <10CBA AB3@1G1 B 1 3C1 A51 83C<A5C4* 31 31 41 4 B1 31 41 *BA)4CC31 <8AB3/1

2F58DC7BCA9ECB235C7BF76C7BC9B6252D6C7B

441<8BC31351A)<31313C1438C131 381B)<41C81 418ABCC45313C1)B8A6A845BC)3C145C1 31 3A )3184BA553 1B)< B3131A5CB8381 31A< 431358313C1)AB 3C1<10CB3C131*BA AB3C/1 43C1 <8B5B<4,1 4C<3C1 33 A<<C1 45C1 3C1 )AB 3C1 <10CB3C1 A5385351 41 <8BBA51 3C1 <8A<8BC1A<B3C13C1)B8A4 3C1?B5B3C131884BA5@1A51A51<31B831<4813C14<<8A13C1  38A)45B3C1 3C1 <8A<8BC1 84B4B33C1 ?B831 6@1 B1 CA51 3C1 <484)B83C1 =35831 >1 41 8CA BA51 31 =4BA51 31 845C<A81 31 <1AA5C1 45C1 31 ) 4531 84BA553 /1 D331 8CA BA51 5A4))351351A)8B31A)< 3,313C18BC1BBB 3131A51B BC3148A8=1B1 3C1385BB83C143453C1 3C1 )1A3C1 31 "A531 D48 A1 B584 3C1 B1 <38)3351 =454 0C381 41 C35CB*B B1 4,1 <484)B83C1 ?35381 351 <B)35C1 4B 31 3C1 3  3C1 A53584BA51 351 *BA)4CC31I@1 )4BC1 4CCB1 41 C35CB*B B1 41 A)4B531B1*A131C81 =A<B)BC4BA51A)8B313C1<1AA*BA8438C/11

43C1 CB54,1 <389C1 >1 =13 31 =531 3  31 B1 84338C31 84<B3)351 3C1 84B35C1 31 )BB831 B)<AC3513183<8C35381A8833)35145C1 3C1)AB 3C1 3C1)45BC)3C1<8B)4B83C131A5338CBA5131 =538B31<1AA5B31351%131<A3AB818381?845C381=3,B4BA5C141C3B513C1<1AAC0CB)3C1 84BA5C1 =A,0A68BA51 C81 3C1 )3)*8453C1 <1AAC051B3C@1 <BC1 381 A< 431 4331 =35C3)* 31 1 )4*A BC)31 A51 =538B31 3C1 A85B31 <481 =4<31 <83531 31 B1 <38)31 31 84 BC381 =4BCBBA513148*A531=4+A3131<1AC<1A8313131C051BC381 41*BA)4CC3/1:3C14<<8A13C1 8BC14B 3C1?)A BC4BA51<481 ,1)4*A B3C@1<333514 A8C1C=4<<0381C81 41A554BCC45313C1 84BA5C131C051BC31 )354B83C/1%138)31 3C1B5B3C1313C184BA5C1<A884B351)D)31D831 )BC3C1 351 83 4BA51 4331 31 5A)31 1 )B8A6A845BC)3/1 51 331 >1 3C1 13 3C1 31 3)<C1 < C1 A53C1 3C1 )45BC)3C1 =44<4BA51 CAC1 35C1 <481 3C1 8 4BA5C1 5B3C1 AB3351 D831 83<8C35C1 45C1 3C1 )AB 3C1 ?/1 J/7@/1 D=3C1 31 4C131 41 <1AA644<4BA51 4<4B1 31 41 3  31 >1 )A 381 CA51 A8B31 <B)354B831 351 A5BA51 31 41 )BB831 =3 31 839AB/1 :31 )D)31 31 0 31 3  4B831B1C=4<<B31C81 =1A8 A31B53853131C813C1353)35C13,3853C1? )BB831BC<A5B*B B1351  )35C158BBC1I@1338414 3)351>138)31D831B58145C1 3C1)AB 3C1<8BBC/151331B 1 B5B1 531 348B4BA51 C0518A531 31 41 4B 31 31 1 A5351 <B)354B831 3C1 3  3C1 B1 CA51 3C1 45BC1  C1 <A81 3C1 )AB 3C1 84B4BC1 ?B)<4451 4B5CB1 3C1 A553C1 =35831 31 41 14B531 31 )A BC4BA5@/1

665947B3B8F7B3CBECCECB

D3C14<<8A13C1) B613 3C1A5BC351>13C1<8A* )4B3C131)A BC4BA51B1CA51<48ABC1 48A8=1B1 5A51 8CA 3C1 451 45C1 31 A)4B531 31 41 <10CB31 ?4  1 3C1 <8A<8BC1 84B4B33C1 31 )B8A4 3C131A8)3C1A)< 3,3C1-13AB817B/1616@113131 41*BA AB31?)3B 3831A)<8135CBA513C1 )45BC)3C1*BA AB3C1>1 =K383115A)3141)4*A BC)3@11351<4CC451<481 41A)<8135CBA51

<10CB31 31 138)A054)B31 31 41 <1AAC051BC31 ?845C381 =3,B4BA5C1 45C1 3C1 453553C1 A< 43C1538B3C141C3B513C1A845B3C13  4B83C@//1

43C1)AB 3C1B58451 =35C3)* 31313C1)45BC)3C1CA518BC1A)< 3,3C1BBB 3C1>14 B*838131 >1454 0C38/14381B BC4BA51<84B313C1 B)B31>1 41CB) 4BA51 41A53<BA5131>1 FA<B)BC4BA513C1 <1AA*BA8438C1 4A8BC451 >1 31 C431 3C1 3)<C1 31 4  C1 A5C1 31 <A81 3C3 C1 31 835C1 CBC1351B535CBB4BA51A511A*35C/1A81 413CBA51A<84BA553 313C1<1AA*BA8438C1<481 A58313C1)AB 3C1< C1CB)< 3C1CA51CA3351B BCC/1 C1CA51< C14B 3C1>14 B*838131531454 0C31 )41)4B31 <38)31 =351 B831 3C1 C84B3C1 31 A58L 31 A1 3C1 C0CB)3C1 31 B45ACB/1 D3C1 4<<8A13C1<38)3351351<48B B381314  3813513)<C183 1 3C1A5BBA5C1A<84AB83C1A<B)4 3C1 <A81 )4,B)BC381 41 <8AB3B1 A1 =34 381 =3A BA51 31 348B4* 3C1  1 ?3538C1 B5843  4B83C1 4B3BC135+0)4B3C1I@1B1531<33351<4C1D831)3C83C1B833)35/1

DA))31 45C1 31 5A)*83,1 483C1 A)4B53C1 3C1 *BA315A AB3C1 =B BC4BA51 31 )AB 3C1 5)8B3C14<4* 3C131<8B83131< C1351< C1B53)351 41<8AB3B1=51BC<ACBB131 831341 C31 584 BC38/1 D3C1 AB C1 <38)38A51 31 84BA554 BC381 31 =A<B)BC381 31 3CB51 31 31 )4,B)BC381 41 <8AB3B145C151A53,31AA1315A)*83,14 4C1 B)4B3C135351>18B831 =3B4B1313C1 C0CB)3C/1:3C18434,1A 4*A84BC135831B55B38C14A)4BB35C1<10CBB35C131*BA ABC3C1338A51 D831CA35C145C131A)4B531351A8133 A<<3)351B C1A5B8A51>13C1<8A8BC1A5CB84* 3C/1

6DEC7BB

1

1 1

7B83161E1123456781E1<1AA84<1B3131 41045A*48B3119A79BCDE942DF4A8CEC1?<B8 B53@/1129BEA81E1 B C84BA5131 41BCBA51=531A531 38A)45B313BCB* 31<481 31)B8A6A845BC)314  31<481 41)1A317::1?7B5B31:B383531B)31:A)4B51G1<4C1313)<C1353B8A5161<BAC3A53@/1E51B1 31 3C14  C1 3C1 <8A<8BC1 84B4B33C1 31 41)B8A64 31 ?A3BB35C1 =4*CA8<BA51 31 31 BCBA51G1 A5BA5131<14C3145 4B83@/1DA)<313513C1B5B3C131884BA5C13C1)B8A4 3C1 =3CC35B3 131 =538B31BC313C1A5353145C15145 3131M#61('N14A8131 41B83BA51B5B353/1

1

1 1

7B831(1E1B) 4BA515)8B3131 =A 3)35145C151C0CB)3131 831310<31843C401 4B1<4815318A31>14*3C/1D3C1CB) 4BA5C1<38)3351=3CB)381 4145B131 )BB831<38931<481 1431)B8A4 3141A8C113)<C/1

1

44ECAC7BB

1

 !"#B $%&B!'(#B $%)%&B *(B+!,'(B %&B $('-B $%B./00/18ABA51 B5C8B3 31 31)B8A4 3C1 31

045A*48B3C/1:45C1E1315B3C131 =55B3818*8B3155A34BA5C1?B5(''@/1 1

1!(#&B$%)%BC9B+('1&B!%B.//2%1441<8ABA5131*BA48*8451 B<BB3143313C1)B8A4 3C1E1

<8A)3CC3C131BC/1B594F282F42B6EA282EBFB3E81('(1?5@1('66(66/1 1 1 1 ,527786ECBB 1

F43C401E1454 1A3381>1B8 4BA5131 B31?)BC1351)A33)351<4815318A31>14*3C@1351A8)3131 O1'1P1?84<<3 4515114)<131A8C3@/1

1 1

"1A3C1 31 "A531 D48 A1 B584 3C1E1 )1A3C1 5)8B3C1 C4BCB3C1 *4C3C1 C81 51 8451 5A)*83131B843C14 4AB83C1?83<8C35451BB1<10CB3)351 3184831311431<1AA51B5B3B3 @/1 43C1 84538C1 <10CB3C1 A*C3834* 3C1 CA51 B3C1 31 =B584BA51 31 AC1 3C1 13)B5C1 31 3B53)35C1A*C383C145C1 313A )3/1

1 1 1 1