Dynamique des populations marines Zaragoza : CIHEAM
Cahiers Options Méditerranéennes; n. 10 1995
pages 53-67
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--- Farrugio H. FAR WEST: An in tern ation al research programme ( 1990-93) for th e N orth western Mediterran ean demersal fish eries. Dynamique des populations marines . Zaragoza : CIHEAM, 1995.
p. 53-67 (Cahiers Options Méditerranéennes; n. 10)
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: (1990-93)
Henri FARRUGIO
IFREMER, 1 rue Jean Vilar,
-
Sete, Francethe have been done at a national
level in the field of assessment and management of
the last decade it has be noted a tendency to implement "analytical" methods of modelling of the exploited stocks, mainly population analysis
models. The techniques of simulation of the
hypothetical of exploitation also in the
These techniques allow to cancel part of the handicaps linked
the stocks che between and use was at a
1990).
the scientists have from the Atlantic that if
some biological exploitation are often
not
known, this should notobstacle to assess a sensitivity
analysis techniques allow to study the impact of such and quantification, in
the of Also, the lack of long of annual
catches and length data should not a applying the
analytical techniques a good idea of the dynamics.
These analyses can which need only few
-
or at leastonly one
-
of sampling of the catches. Finally it that it was possible to perform VPA on ages kngah of some meditenanean stocks exploited with gears aad to analyse theSome mehdobgical on sampling also
been donc previously. of such that it was possible to obtain, via a sustained action in time, good data about the fishing activities ni the
This conceptual evolution have led in 1989 to the setting up of a
an attempt to solve the question of the of the &ta collecting and at a 1990 to 1993, a
Spain scientists belonging to 8
of the mast of the to the Tuscan
(fig 1).
This 4 (known the code
was aimed at the out of a common methodology the study of some
of the most topics in the Spanish and following
achieved the 4 of
-
the setting up of a common &ta base on catches, efforts and biological information,-
the improvement of the knowledge of demographic and biological characteristics-
a general reflexion about the methods and technics to be created a 4 adapted fitture some demersal andpelagic fuh sto& of major economic value,common assessments and managements of the western mediterranean jìsheries.
-
an improvement of the knowledge biological parameters, fleets dynamics, fishing-
an approach of the modelling of some of the most characteristic problems regarding the-
some preliminary predictive studies about the effects of changes in fuhing patterns-
an improvement of the knowledge about economical characteristics of the fisheries efforts and capacities,demersal fiheries the countries associated in the project,
(assessment of stocks, sensitivity analysis, simulation of selectivity).
sector.
The objective is
et a1.,1991,1994) and in
1.
of the
of the
The final two
main kinds of files :
-
a common wded fleets,files can also of
-
contain and biologicalsamplings. Some of them "exhaustive data files" aimed at the stocking of the global catches and fishing efforfs, and of
of of of "sampling data
at the stocking of non equipments of the boats, catch samples
of the in 3+
on The
scheme of of the base been built using the
fundamental concepts
The has used tool a basis the
biological and explaitation aspects.
collection of at
caught by the hake merluccius), sole
(Solea vulgaris), sea bass l a b r a ) , (Sparus auratus) and
k k t e u s antennatus. it is one
of of
The been
of 1990 to the end of to the seasons and the species, the biological monthly samplings took place in
board of
also
which where carried in the of of the
audies.
The data base been also not) of
in
of dynamics, the data base has also been fed with
.of fleets
of stock assessments, essentially by using
example the and ANACO models
cohort of exploited stocks,
aimed at
Fig. 1. programme geographic coverage
of thc studied.
sanlpling points
0
di Scicnzc dell'Ambicnte e dcl di
Grupo dc dc dc Cicncias
dc Français dc dc la
di Zoologia d i Gcnova
Arcas cikd in the text : a: sca b: gulf of c: Alicanlc d: Valencia e: Cataluña sea' g: gulf of Lions h: sea i: sea.
veral computer programs have been set up, aimed firstly at the facilitation of stocking and sorting operations and secondly at the interfacing between the data base and the assessment modelling programs (mainly the one).
The complete process, from basic data to assessment results can schematized as shown in fig. 2 below :
- > { F I
errors
I <
I?
1
>
I I
l-
II
I
f o r
I -
-
2.1 Growth of merluccius)
it is well known that
of
usable pieces) is This
of of the
of to
to
stock assessments. The of this of
existing computation models to the available Sets of &ta and to
k e n done:
-
the same set of data(same arca),
-
ofareas).
where by sexes,
sampled at landing in
-
Splitting in of a smoothing onin er al.,
the et al., 1988), using inputs these
2
-
Assessment of the same sets of data,1 of
-
et al., 1990)1991 data.
-
et al., forin the Gulf of the
used estimahg paramedess. of
she growth rates of of life.
A model also been established, taking into account sizes
of some 20 for 1 40 cm
than 50 of life females, and maximum sizes 110
discounting the concept of the existence of a hake.
also been applied to
(Alicante, Spain). The by
vessels al Santa
application of
on files,
of the VBGF
Of the
to goWh
of
& 1979) methods the analysis of
cm hasized on the identification of annual is a goal t0
S
of57
2.1.2.
Gulf of to than one
could think to
that mean is 16 cm at the end of the of life. The
The of 1.18 obtained by means of modal length- high estimates, showing the
The of
Finally, a set of
of the spccies, the fitting of to
conditions of of to of the daily
otolilhs
For 1, 0.4 have h e n
as convenient; the global stock assessments i t has to use the sexes; for the Guif of it is best
f i t
and to values which do a suitabk
on thesc of the Von
to be used as population dynamics analyses on the of
S p a n , z i g u r .
.
TTyrrhen.
both sexes
I I
.48 49.8 - 2 8
-.O44
emales es
k t o k t o
5 5 . 0
Tab. 1.- , Span.Med = Spanish = Gulf of Lions,
f.igur..p=
sea continental shelf, Ligur.S =sea continental slope,
Ligur.T= total, Tynhen. = Tyrrhenian22. Growth of (Arkfew anfennutus)
2.2.1. and methods
of of
is to the change of
moulting. So
techniques to of
ages. Size re on the catches have been
splitted by sex and weightcd by
f
catches in...). of and
2.2.2.
by a consensus have
been met on a set of to be used
stock assessments on this species (tab. 2).
es
O P GENOA
~
Tab. 2.- clristeus antennatus
2.3 OF
CASE OF to attempt the modelling of
l
2.3.1.
As of not long enough to
achieve classic of the case-studies
So the use of the model and has
been adopted. This modcl was specially built by the to
by ages
or state on so
that it is not suitable of these
a! the moment).
From the catch
(VPA)
tools of he can
VPA in exrensu, based on (F)
analysis of the of
a change of due to changes in
applicable to the study of
The also sense
may be can be size to
age The outputs of file to be onto a
l of gears.
59
k e n use on
sets of data have been used in to
of assessments based on lenght
have been done listed in table 2.
I I
G u l f
&
+ +
BASS SEA
G u l f G u l f G u l f o f
2.3.2.1.
Using the data on period 1988-1992, an analysis
Y/R) of the Spanish "hvant" in
of state of this stock to be
an exploitation scheme with
subsisting
This is
of the Y/R
to was
of
of this stock.
at of the
and models have to this 13
significant between the by the two methods.
For the of
surmuletus)
of
1980
-
1991, have been used two specificpseudocohorts on which VPA and Yield per analyses were performed
with
the60
inadequacy of the is applied. A
shown that using a (60mm) will in a significant of both
spccics yields and biomasses.
in a of and
populations status has been done using VPA and calculations. This
used the data catches on islands.
Exploitation pattern lightly one from The mean lengths and ages
in level is
assessments have shown a biological all
A on thc sea
coming both 13 out 1990 to 1993 and samplings at
landing. nlodcl on (1990,91,92) and on an
this The two to the same about
the state of this stock (i.e. hcavy exploitation). was shown that of the
rcsufrs by taking into account the bio-ecological
obscwations which h a w bccn donc on the last 25
basin the 1992
hakc by of uscd by the been uscd
to somc VPA and to one
have shown a heavily biological with a low age
and a values of the two bccn
shown, but in both a s e s low if to the
A of of hake in two non-adjacent of
Cataluna, Spain) has been performed
using the 1990-1992 these two
computations have been done both on lengths and ages
of in the
project The to
and Catalan that the hake can be
as the two
the Gulf of Lions, the exploitation of the hake stock is and typcs of Some analyses of its dynamics have bccn
and to the biological
was studied, as well as the These analyses have shown that, a biologicaí point of view, this stock can as heavily exploited. Another
is the evidcncc t h d mainly small individuals (i.e. have a gears
-
such as gillnets and longlines-
An assessment of the same stock has been 1988-1991, using two methods of population analysis based on
The of been a methodolgical point of
ones to the level of exploitation of hake. The
two methods give quite similar rcsdts and mean biomass. The
fishing martaiity mte on juveniles is
and for the biomass/mcan an level
of exploitation but not so high to bc 2.3.2.2.
-which is only fished at sea- the sea bass, the sea
of the main species of small-scale and
fleets in the Gulf of Lions. The on the 1986, 1987, 1991 and 1992 catch
to thcsc activitics to build a diagnosis of
61
the stocks of
period,
on and on the possibleof an of the of of fleets.
Thc havc shown that thc stocks can bc as biologically hcavily cxploitcd. They stay at low bionlass lcvcls but is to any
of Thc most of the to the
which hcavity on thc small-scaìe
of thc resulted in in yield
the fleets.
The evolution of the fishing of Aristeus antennutus on the Catalan
coast has also of The exploited
as wcll the mean size of
1983-1991; to this species has
in
of 0.5 fcmalcs and 0.8
This shows that thc fishing on this spccics is closc to the optimum.
A of A . utrfetrnatus Catalonia and Valcncia has also bccn donc.
An annua1 of by analyscd a 6 (1983-1989)
using LCA and Sonx rtnalyscs in to gct a of the
status of the stock. Scnsitivity of K A and
studicd in to evaluate the cffcct of The stocks
not t o bc far from the stcady state and a
hcncc it was possible to use LCA. The vs level closc to
not they yield.
The status of Gamba by in
adjacent zones and been
using VPA and analyses. The that is close to the
biological not show evidence of
2.3.2.3. of the stocks
l
The pol?ulatjon
aimed at the of the feasibility of a to analytical
stock assessments in
beyond this some
about the status of some stocks a biological
point of view we can state that we
diagnoses of biological full exploitation, This
situation seems to be the of both the conditions of the exploited populations and the to them. limitatcs today's catchcs
to WC
that if to a (and one) was
on thc past, such an
into a of biological of The situation
Seems to to at the
moment.
2.4. Studies gear selectivity
bcen achieved to obtain additional data which on the possible cffccts of changes in mesh sizes on the age composition of the stocks. The data of
of sea have some
among which have to be noticed the use of
of the these mixed
fisheries it can bc cxpcctcd that
62
studies on by
new aimed at selectivity evaluations by the
method, took place on local fishing boats in
(Spain) between and The data collcctcd, especially on
hake, of
and cumes assessed.
of bcen to compile all these
and to discuss about the methodologics, the of the
of on
3.
The of an
to in thc economical field to
This has been done stating that the methodological fundamcnts of totally applicable to
basic objective of this of the was not to analyscs but to imagine
a databaqe which will contain the clcmcnts to establish the managcmcnt
at the main efficiency all of the systcm". fact, it to a whole, bccause fishing is linked to
take an placc.
to the main of the database which includcs mainly eco-biological and of the files which could the economical of the bcen defincd. They consist mainly in two types of and,fleets and socio-cconomical data. of this (especially the technical one) was yct includcd in of
common database but at a step of
easily be eliminated, as the between do not poscs any technical
a encoding
of of the socio-
economical This methodology has been tcstcd on the Andaluz fishing the files
have been fed and data and sonic cxamplcs of
have bcen 4.
of b c c n u c to analyzc thc of
the hakc unit of of in two of thc
the Gulf of basin. in
the hake cpue (kg/day) have bcen to be thc of thc combination of thc following (hp), fishing fishing zonc and tinlc of (month). With
to a distinction was t o thc opening of the mouth.
into two and to d c t c n i n c thc
of the cpue a modcl has bcen used.
The daily mcan catch, vcsscl and month, showcd a high with to thc to of this study, the expcctcd hakc yicld vcsscl mainly dcpcnds on the time of and fishing zone and, to a on the vcsscl
of
withthe mcntionncd,
of hake in this area has been
in and
have been of abundance of
a of the it can be some
conclusions, that
of out up to now,
of the dynamics of exploited populations, does not
allows to to of the national bodies.
be based on full knowledge of the system and shouid
out concept.
of one of the
in of the typology of
lead to the dcfinition of well individualized units of exploitation and managemcnt.
Thc of thcsc analyses and an of the
quality of the statistics in a of species, an
of thc knowlcdgc on the biological (age, on the
Studies on
and also to be as well as thc
of the mechanisms of biomosscs fluctuations in space and time. Still at thc fundamental lcvel, tbc
to explain the to and
those of the fleets' dynamics bccome of too.
such a of
databases will be of a of the situation as a whole.
a socio-cconomic point of of the
evolution of the investments and the capital, of seems indispensable.
The also
-
extending-
ofand of only
sketchy knowledge is available.
64
FARRUGIO, H., F. ALVAREZ, J. LLEONART and DE RANIERI.- 1991. Study for assessment and management of fisheries in the western mediterranean, part 1. €€C-FAR MA 1-232 Final report 454 p.
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GAYANILO ir, F.C., M. SORIANO and D. PAULY.
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1988. A draft guide to the compleat ELEFAN.Software2, ICLARM, p.
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1989. Computer programs for fish stock assessment. ANACO : Software for the analysis of catch data by age group on IBM PC and compatibles. FAO Fish. Tech. Pap., 101 (Suppl. 3): 73 PP.SAILA, and PRAGER.
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CASTILLO CABEZAS M., RUIZ BRAVO G. & RUIZ MOLINA A.- Management of fisheries:
CASTILLO CABEZAS M., RUIZ BRAVO G. & RUIZ MOUNAA- Socioeconomic information system DEMESTRE M., LLEONART J.- 1994. Population dy'hamics of Aristeus antennatus (decapoda:
proposal for a methodology of investigation. id.:
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dendrobmchiata) in the northwestern mediterranean. id.:
DEMESTRE M., CARBONELL A., M A R T I N E Z - W P.- Analysis of the exploited population of Aristeus antennatus (Risso, in three adjacent zones n the Spanish Mediterranean. id.:
DEMESTRE M., MARTIN Optimum exploitation of a demersal resource in the Western Mediterranean: the fishery of the deep water shrimp Aristeus antennatus (Risso, id.:
DEMESTRE M., CARBONELL k- Growth of the shrimp Aristeus antennarus: results of the FARWEST programme working group. id.:
FARRUGIO LE CORRE VAUD0 Population dynamics of sea bass, sea bream and sole exploited the French multigears demersal fishery in the Gulf of Lions (Northwesterm Meditemmar$. id.:
GARCIA-RODRIGUE2 Determination of the growth parameters using length frequencies:
application to the Mediterranean hake (Merluccius merluccius) Santa Pola bay (Spain). id.:
GARCIA-RODRIGUE2 M., RECASENS L,- Assessment of the hake (Merluccius merluccius) population in the Levante zone (Murcia-Valencia) of the Spanish Mediterranean area. id.:
GIL-DE-SOLA SIMARRO L.- Trawl selectivity in the Western Mediterranean and Adriatic sea fisheries. id.:
GIL-DE-SOU SIMARRO GAZA M.- Distribution of hake ,(Merluccius merluccius) in the Alborran sea. id.: 250-257.
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MARTIN P., VIVA C.- Analysis of the variations of the Merluccius merluccius catch per unit of effort (cpue) in two areas of the Northwestern Mediterranean. id.:
MORALES-NIN B., ALDEBERT Y.- Growth and birthdate distribution of juvenile Merluccius merluccius in the Gulf of Lions (NW Mediterranean) determined by otolith microstructure and length frequency distribution. id.:
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66
OUVER P.- Catch fluctuations in a trawl fishery analysis of fluctuations observed in the trawl fleet OUVER P., REilONES MASSUTI E.- Analysis of the exploitation of hake in Majorca id.:
OUVER P., MASSUTI E., REirONES A cornDarison of results obtained bv mean a cohort landings of the Balearic islands. id.:
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management of the FARWEST database files. id.:
different types of trawl in the Northern Tyrrhenian sea. id.:
(Merluccius merluccius) in the Spanish Mediterranean fisheries. id.:
67