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Plicathyridine brachiopods from the Frasnian (Upper Devonian) of the Middle East and Belgium

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S27B - ISOTOPIC SIGNATURES FOR THE HABITAT AND GROWTH OF THE MIDDLE JURASSIC AMMONOID QUENSTEDTOCERAS

Moriya, Kazuyoshi1, Hirano, Hiromichi1, Oba, Tadamichi2, Tanabe, Kazushige3and Kulicki, Cyprian4

1 Department of Earth Sciences, Waseda University, 1-6-1 Nishiwaseda, Shinjuku-ku, Tokyo 169-8050, Japan 2 Graduate School of Environmental Science, Hokkaido University, Kita-10 Nishi-5, Kita-ku,

Sapporo 060-0810, Japan

3 Department of Earth and Planetary Science, the University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo 113-0033, Japan

4 Institute of Paleobiology, Polish Academy of Science, ul. Twarda 51/55, PL− 00−818 Warsaw, Poland

e-mail: kmoriya@aoni.waseda.jp

Oxygen and carbon isotopic compositions are presented for extremely well preserved shells of the Callovian (Middle Jurassic) ammonoid Quenstedtoceras recovered from Lukow, western Poland. Averaged oxygen isotopic temperatures of two individuals analyzed (39 and 31 mm in maximum diameter, respectively) provide palaeotemperatures of 15 and 17 °C. Considering that the oxygen isotopic temperatures of

Gryphaea (bivalve) and nektonic vertebrate remains recovered from the Callovian Oxford Clay of the UK

show 11 and 24°C, respectively (Anderson et al, 1994; Price and Page, 2008), these two Quenstedtoceras individuals seem to have been nektonic swimmers within the Jurassic water column. While the oxygen isotopic compositions of these two individuals do not show a distinct systematic trend, carbon isotopic compositions (δ13C) display a sigmoidal fluctuation in both individuals. Although the amplitude of δ13C variation is different, one period of isotopic fluctuation corresponds to three quarters to one whorl of growth in each individual. If we assume that the δ13C of the specimens analyzed reflects the δ13C of dissolved

inorganic carbon (DIC) in the water column where these specimens lived, and that the fluctuation in δ13C of DIC is annual, we can estimate the growth rate of Quenstedtoceras as three quarters to one whorl a year.

S7 - PLICATHYRIDINE BRACHIOPODS FROM THE FRASNIAN (UPPER DEVONIAN) OF THE MIDDLE EAST AND BELGIUM

Mottequin, Bernard1and Brice, Denise2

1Service de Paléontologie animale et humaine, Université de Liège, Bât. B18, allée du 6 Août, 4000 Liège, Belgium

2Faculté Libre des Sciences et Technologies, Université Catholique de Lille, 41 rue du Port, 59046 Lille Cedex, France

e-mail: bmottequin@ulg.ac.be

Plicathyridines (Athyridida), which disappeared during the Late Frasnian biological crisis, are well-known in the Frasnian of the Russian Platform and the Altai-Sayan Province, but these distinctive brachiopods also occur in Frasnian-aged rocks of the Middle East (Iran and Afghanistan) and Western Europe (e.g. Belgium). In Iran (Esfahan Province), specimens tentatively assigned to Plicathyris are present in a biostromal horizon apparently located just below the Palmatolepis jamieae conodont Zone whereas similar specimens occur in east-central Afghanistan (western part of the Dacht-e-Nawar). In this area, they are associated with an unidentified species of Cariniferella probably belonging to the C. tioga group. These orthides may indicate the P. punctata Zone in comparison with their occurrence in the Frasnian succession of the Boulonnais (northern France), but further research is needed. In southern Belgium (Dinant Synclinorium), plicathyridines are present only in the shaly La Prée (Nismes Formation) and Ermitage (Moulin Liénaux Formation)

members (P. falsiovalis to P. punctata conodont zones) and belong to the genus Anathyris (A. helmerseni group); they are thus absent in the Upper Frasnian of the Namur-Dinant Basin. The plicathyridines constitute only a minor part of the Frasnian brachiopod assemblages recognized in the three studied areas.

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