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Mise au point sur les méthodes de dépistage des parasitoses chez les bovins (autopsies exclues)

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Open

Archive TOULOUSE Archive Ouverte (OATAO)

OATAO is an open access repository that collects the work of Toulouse researchers and makes it freely available over the web where possible.

This is an author-deposited version published in : http://oatao.univ-toulouse.fr/ Eprints ID : 8628

To cite this version :

Fourcade, Raphaël. Mise au point sur les méthodes de dépistage

des parasitoses chez les bovins (autopsies exclues). Thèse

d'exercice, Ecole Nationale Vétérinaire de Toulouse - ENVT, 2012, 169 p.

Any correspondance concerning this service should be sent to the repository administrator: staff-oatao@inp-toulouse.fr.

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ANNEE 2012 THESE : 2012 – TOU 3 – 4068

MISE AU POINT SUR LES METHODES DE

DEPISTAGE DES PARASITOSES CHEZ LES

BOVINS (AUTOPSIES EXCLUES)

BBBBBBBBBBBBBBBBB

THESE

pour obtenir le grade de DOCTEUR VETERINAIRE

DIPLOME D’ETAT

présentée et soutenue publiquement devant l’Université Paul-Sabatier de Toulouse

par

FOURCADE Raphaël

Né, le 3 Février 1986 à NANCY (54)

___________

Directeur de thèse : M. Philippe DORCHIES

___________ JURY PRESIDENT : M. Alexis VALENTIN ASSESSEURS : M. Philippe DORCHIES M. Philippe JACQUIET

Professeur à l’Université Paul-Sabatier de TOULOUSE

Professeur à l’Ecole Nationale Vétérinaire de TOULOUSE Professeur à l’Ecole Nationale Vétérinaire de TOULOUSE

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Ministère de l'Agriculture et de la Pêche ECOLE NATIONALE VETERINAIRE DE TOULOUSE

Directeur : M. A. MILON

Directeurs honoraires M. G. VAN HAVERBEKE. M. P. DESNOYERS 3URIHVVHXUVKRQRUDLUHV :

M. L. FALIU M. J. CHANTAL M. BODIN ROZAT DE MENDRES NEGRE

M. C. LABIE M. JF. GUELFI M. DORCHIES M. C. PAVAUX M. EECKHOUTTE

M. F. LESCURE M. D.GRIESS M. A. RICO M. CABANIE M. A. CAZIEUX M. DARRE

Mme V. BURGAT M. HENROTEAUX

PROFESSEURS CLASSE EXCEPTIONNELLE

M. AUTEFAGE André, Pathologie chirurgicale

M. BRAUN Jean-Pierre, Physique et Chimie biologiques et médicales

M. CORPET Denis, Science de l'Aliment et Technologies dans les Industries agro-alimentaires

M. ENJALBERT Francis, Alimentation

M. EUZEBY Jean, Pathologie générale, Microbiologie, Immunologie

M. FRANC Michel, Parasitologie et Maladies parasitaires

M. MARTINEAU Guy, Pathologie médicale du Bétail et des Animaux de Basse-cour

M. PETIT Claude, Pharmacie et Toxicologie

M. REGNIER Alain, Physiopathologie oculaire

M. SAUTET Jean, Anatomie

M. TOUTAIN Pierre-Louis, Physiologie et Thérapeutique

PROFESSEURS 1° CLASSE

M. BERTHELOT Xavier, Pathologie de la Reproduction

Mme CLAUW Martine, Pharmacie-Toxicologie

M. CONCORDET Didier, Mathématiques, Statistiques, Modélisation

M DELVERDIER Maxence, Anatomie Pathologique

M. SCHELCHER François, Pathologie médicale du Bétail et des Animaux de Basse-cour

PROFESSEURS 2° CLASSE

Mme BENARD Geneviève, Hygiène et Industrie des Denrées alimentaires d'Origine animale M. BOUSQUET-MELOU Alain, Physiologie et Thérapeutique

Mme CHASTANT-MAILLARD Sylvie, Pathologie de la Reproduction M. DUCOS Alain, Zootechnie

M. DUCOS DE LAHITTE Jacques, Parasitologie et Maladies parasitaires

M. FOUCRAS Gilles, Pathologie des ruminants

Mme GAYRARD-TROY Véronique, Physiologie de la Reproduction, Endocrinologie M. GUERRE Philippe, Pharmacie et Toxicologie

Mme HAGEN-PICARD Nicole, Pathologie de la Reproduction M. JACQUIET Philippe, Parasitologie et Maladies Parasitaires

M. LEFEBVRE Hervé, Physiologie et Thérapeutique

M. LIGNEREUX Yves, Anatomie

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M. SANS Pierre, Productions animales

Mme TRUMEL Catherine, Pathologie médicale des Equidés et Carnivores

PROFESSEURS CERTIFIES DE L'ENSEIGNEMENT AGRICOLE

Mme MICHAUD Françoise, Professeur d'Anglais M SEVERAC Benoît, Professeur d'Anglais

MAITRES DE CONFERENCES HORS CLASSE

M. BAILLY Jean-Denis, Hygiène et Industrie des Denrées alimentaires d'Origine animale

M. BERGONIER Dominique, Pathologie de la Reproduction

Mle BOULLIER Séverine, Immunologie générale et médicale

Mme BOURGES-ABELLA Nathalie, Histologie, Anatomie pathologique

M. BRUGERE Hubert, Hygiène et Industrie des Denrées alimentaires d'Origine animale

Mle DIQUELOU Armelle, Pathologie médicale des Equidés et des Carnivores

M. JOUGLAR Jean-Yves, Pathologie médicale du Bétail et des Animaux de Basse-cour

M MEYER Gilles, Pathologie des ruminants.

Mme LETRON-RAYMOND Isabelle, Anatomie pathologique

MAITRES DE CONFERENCES (classe normale)

M. ASIMUS Erik, Pathologie chirurgicale

Mme BENNIS-BRET Lydie, Physique et Chimie biologiques et médicales M. BERTAGNOLI Stéphane, Pathologie infectieuse

Mle BIBBAL Delphine, Hygiène et Industrie des Denrées alimentaires d'Origine animale

Mme BOUCLAINVILLE-CAMUS Christelle, Biologie cellulaire et moléculaire Mle CADIERGUES Marie-Christine, Dermatologie

M. CONCHOU Fabrice, Imagerie médicale

M. CORBIERE Fabien, Pathologie des ruminants

M. CUEVAS RAMOS Gabriel, Chirurgie Equine

M. DOSSIN Olivier, Pathologie médicale des Equidés et des Carnivores

Mlle FERRAN Aude, Physiologie

M. GUERIN Jean-Luc, Elevage et Santé avicoles et cunicoles

M. JAEG Jean-Philippe, Pharmacie et Toxicologie

Mle LACROUX Caroline, Anatomie Pathologique des animaux de rente

M. LIENARD Emmanuel, Parasitologie et maladies parasitaires M. LYAZRHI Faouzi, Statistiques biologiques et Mathématiques

M. MAILLARD Renaud, Pathologie des Ruminants M. MATHON Didier, Pathologie chirurgicale

Mme MEYNAUD-COLLARD Patricia, Pathologie Chirurgicale M. MOGICATO Giovanni, Anatomie, Imagerie médicale

M. NOUVEL Laurent, Pathologie de la reproduction

Mle PALIERNE Sophie, Chirurgie des animaux de compagnie

Mme PRIYMENKO Nathalie, Alimentation

Mme TROEGELER-MEYNADIER Annabelle, Alimentation

M. VOLMER Romain, Microbiologie et Infectiologie (disponibilité à cpt du 01/09/10)

M. VERWAERDE Patrick, Anesthésie, Réanimation

MAITRES DE CONFERENCES et AGENTS CONTRACTUELS

M. BOURRET Vincent, Microbiologie et infectiologie

M. DASTE Thomas, Urgences-soins intensifs

ASSISTANTS D'ENSEIGNEMENT ET DE RECHERCHE CONTRACTUELS

Mlle DEVIERS Alexandra, Anatomie-Imagerie M. DOUET Jean-Yves, Ophtalmologie Mlle LAVOUE Rachel, Médecine Interne Mlle PASTOR Mélanie, Médecine Interne M. RABOISSON Didier, Productions animales

Mle TREVENNEC Karen, Epidémiologie, gestion de la santé des élevages avicoles et porcins

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'''0'=4#%&2'1"2'(+%2Q1&'.&'G%I4(<'&('.&'igW3'

)"4+' (*' /*,,%"2' .&,' +<41,' *++*2$D,3' .&,' ,8<D1*,' &2' H<Q##&,' .&' G%"' &(' .&' $D"#3' &(' .4' ?+*2\*%,:' )"4+' (&,' 8<&6&4@' ?#*1G"B*2(,3' >4%' 7&' #5&,/;+&' ?"2(' #&4+' /&(%(' &??&(' ,4+' #&,' 7*/"2*%,&,:'

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'''0'=G3'1"2'*4(+&'(+%2Q1&'&2'G%I4(<3'&('*1%'.&/4%,'#&'8"##;$&3'

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(11)

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'''0' ("4(&' #*' 8#%>4&' .&' G%I4(<,'Z' T*4+*3' U*4/%&3' P"+"3' J&23' =4743' -*+%&3' V&#/<%2&3' &(' )&(%(' d"+1*(3'

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d+*2\"%,'`"4+"4G#&3'/"4+',&,'8"2,&%#,'74.%8%&4@:' ' ' '''04@'.%??D+&2(,'8*G%2&(,'6D(D+%2*%+&,'>4%'15"2('*884&%##%'#"+,'.&',(*$&'"4'.&'/+"/<B#*@%&:' ' ' '''0'("4(&,'#&,'/&+,"22&,'*B*2('/*+(%8%/D'9'#*'8"#"2%&'C"#6*B'9'V"4*+2&2&I3'$+R8&'9'>4%'75*%' /*,,D'.&'?"+1%.*G#&,'1"%,'.&'74%##&('.&/4%,'WXXj3'&('/#4,'/*+(%84#%;+&1&2('Z'' '

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(12)

! !!" '''0'#5D8"#&'6D("3' ' '''U"4('.5*G"+.'9'1&,'/#4,'/+"8<&,'*1%,3'("4,'/#4,'#"4+.,'#&,'42,'>4&'#&,'*4(+&,3'1*%,',*2,' >4%'8&,'8%2>'*22D&,'25*4+*%&2('/*,'D(D'*4,,%'+%8<&,'&('+*??%2D&,'Z'' ' '''0'^&+$3''

k4&' .&' 8<&1%2' /*+8"4+4' .&/4%,' 8&' ?*1&4@' /+&1%&+' (.' .&' 1*(<,' &2' G%I4(<3' &(' 8&' ?"4' +%+&' .542&' <&4+&' >4%' ,5&2' ,4%6%(M' 0' (&,' 84%(&,' &2' /+D/*' ,&' ?%2%,,*2(' 2D8&,,*%+&1&2(' 1*#3' 9' #5%2,8+%/(%"2'*4'("4+2"%'.&'?""('&2'fgW'*/+;,',5A(+&'?*%('6%+&+'.&'8"4+,'.&'/<B,%>4&:')4%,'6l2(' #&' (&1/,' .&' #5D8"#&:' 0' ("4,' 2",' ?"4,' +%+&,' &2' G"41' "4' &2' 8"4+,g(.g8#%2%>4&:' 0' ("4(&,' 8&,' ,"%+D&,' 1D1"+*G#&,:' 0' 2",' G"%,,"2,' /%D$D&,3' 2",' ?#*>4&,' 8"2(+&' #&' G*+3' 2",' 8"4/,' .&' /%&.,' .*2,'#&,'6&++&,3'&('2",'2DG4#%,*(%"2,'.&'G%;+&:'0'2"(+&'e&&H'&2.'^US'8"++;I&M'0'(&,'.*2,&,' ?*\"2' ?"++&,(' $41/3' (&,' 2"1G+&4,&,' 8<"/&,' 9' #*' +D8"28%#5' .&' /"4#"(' E+*//&##&' ("%3' 02*K,' m&+G&+(MF3' ("2' *88&2(' 1&+.%>4&' &(' ['*4']' 8"%??&4+:' 0' ("4,' 2",' ,*1&.%,' .&' 1"%,%,' */+;,' #&,' G"41,3' 9' *$"2%,&+' *4' 8%2D' "4' *4' -*8' V":' 0' ("4,' 8&,' */D+",' *6&8' `<%15,' 8"11&' (+"%,' #"",&+,:')"4+'(*'$&2(%##&,,&3',B1/*(<%&3'&('8"4+("%,%&'*6&8'#&,'$&2,'&2'G"41:'0'(*'8*+*G%2&'9' /#"1G,'&2'G"6%2&'&('*4'(%+'*4@'/%$&"2,:' ' ' '''0'`<%15,3'' )"4+'(&,'?*84#(D,'<"+,O2"+1&,'.&'8"1/+D<&2,%"23'(*'(A(&'.&'14#&'E(4'2&',&+*%,'/*,'G*,>4&' /*+'<*,*+.'nF3'(*'("#D+*28&3'(*'?%2&,,&'&2'G"41'&('*6&8'#&,'$&2,'>4&'(4'2&'8"22*%,'/*,:'0'(*' +D,%,(*28&' *#8""#%>4&' *4' +%8*+.' #&' /+&1%&+' ,"%+' .&,' oLpLU' WXXq3' &(' (*' ,4G%(&' /*,,%"2' .&,' 1;8<&,:')"4+'("2'*1"4+'.&,'8<%&2,'G"l(&4@'&2'/#*,(%>4&'&('/"4+',%??#&+'8"11&'42'+",,%$2"#:' 0' (&,' (*#&2(,' .&' 8<*2(&4+' 8"224,3' +&8"224,3' &(' *//+D8%D,' .&' ("4,:' )"4+' 2"(+&' *1"4+' 8"1142'.&'#*'#*2$4&'.&'C<*H&,/&*+&'E6%6&'#&'$+"4/&'Yh'rF:')"4+'A(+&'42'$+",'$&&H'.&,'7&4@' 6%.D",',4+'sOJ"@'ihX:')"4+'*%1&+'#&,'B*"4+(,'G4#$*+&,:' ' ' '''04'J&#$&3'' )"4+'(*'8*/*8%(D'9',"+(%+'*4(*2('.&'8"22&+%&,'&2'42&'1%24(&:')"4+'("2'*.+&,,&'#D$&2.*%+&3' (&,'1"K,&,3'&('(&,'"+(&%#,'+&8(%#%$2&,:'0'(*'6%,%"2'(+;,'/&+,"22&##&'.&'#*'1".&:'0'(&,',/+*B,'ihX' &(' (&,' ."4#&4+,' %2(&,(%2*#&,:' 0' 2",' >4*+(,' .5<&4+&' 8#4GG%2$' &2' G"41' &(' 2",' 8"28"4+,' .&' 1"48<&:' 0' (&,' (*#&2(,' .&' G+&*H' .*28&+:' 0' (&,' G#*$4&,' &(' 7&4@' .&' 1"(,' /"4++%,' *6&8' #&,' /+D2"1,3' /"HD1"2,3' &(' *4(+&,' *2%1*4@3' >4%' 2&' 1&' ?"2(' /*,' +%+&:' 0' (*' 8%++<",&' *6*28D&' &(' ("2'/+"$2*(<%,1&:')"4+'A(+&'42'.4+'9'84%+&3'&('*6"%+'/+%,'#&',(*(4('.&'#&*.&+:'0'(&,'.&4@'/#4,' $+*2.&,'/*,,%"2,'Z'(&'?*%+&'6%+&+'.&'#5*1%8*#&'.&,'D#;6&,3'&('#*'.&2(%,(&+%&'D>4%2&:'

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'''0'J*#*3''

)"4+' 2&' /*,' 6"4#"%+' *.1&((+&' >45%#' 25B' *' /*,' .&' ['$']' 9' #*' ?%2' .&' ['/*%2']' "4' ['8"/*%2']3' &(' /"4+'/*+(*$&+'#*'/*,,%"2'.&'#5&@8#4,%"2'.&'#5*1%8*#&'.&,'D#;6&,'*6&8'#&'J&#$&:'0'(*'8*/*8%(D'9' (&' ,&+6%+' .&' ("2' ?+"2(' /"4+' "46+%+' #&,' 2"%,&((&,3' #&,' 2"%@3' #&,' /"(,' .&' B*"4+(,3' #&,' G"l(&,' .&' 8"2,&+6&3' #&,' #*1/*.*%+&,3' #&,' &@(D+%&4+,M' 0' (*' (A(&' .&' -%8H&B' &(' (&,' #42&((&,' ?#%8O?#*8' %28*,,*G#&,:'0'(*',*#1"2&##",&'&('("2'<&+/&,:'0'(&,'2"1G+&4,&,'8"2>4A(&,'8"224&,'&(',"4,O 1*+%2,3'*%2,%'>459'(&,'*12D,%&,'/",(OG"41'E<&4+&4,&1&2('>45%#'B'*'#5"%,&*4'.&'-DI*+.FM' '

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(13)

! !#" '''0'd+*2I"4%##&3'' )"4+'("2'*1"4+'.&'#*'$*,(+"2"1%&'?+*2\*%,&'Z',*48%,,"2'&('/%2*+.3'&('.&,'/&(%(,'/#*%,%+,'.&' #*'6%&'Z'8<*,,&3'/A8<&'&('G%(4+&:'L$*#&1&2('/"4+'(*'/*,,%"2'.&'#5&@8#4,%"23'.&,'8"4/,'.&'(A(&3' &('.&,'&@(D+%&4+,'E.D8%.D1&2(3'>4&'.&'/"%2(,3'"4'/"%2$,3'8"1142,MF:'0'(&,'$+*2.,'(*#&2(,' .&'2*$&4+'&('(&,'/+"$+;,'*4'/%2$'/"2$3'$+R8&'*4'/%8"#"O/"2$:'0'("2'*1"4+'.&,'+*//"+(,'.&' ,(*$&'&2'0W3'&('.5*4("/,%&,'&2'0i:'0'2"(+&'+D$%1&'*2$#*%,'/&2.*2('.&4@',&1*%2&,'Z'>4*(+&' /%2(&,'.&'G%;+&,'&('8<%/,'.&6*2('#*'(D#D'#&',"%+:'0'2"(+&',&6+*$&'&2'=*8H'V*2%&#5,:'0'(*'/*,,%"2' .4'=t$&+1&%,(&+'8<*4.'&2'/#&%2',"#&%#'*4'G"+.'.&'#*'/%,8%2&:' ' ' '''0'S%28&3'

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'''0'`B+%##&3''

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'''0'`*13''

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'''0'`<5(%('d*6+&*43'

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(14)

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'''0'^"1*+3'

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'''0'`%#43'''

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'''04@'P&2%?#"+,'Z')&/5I'E['B'*'/*,'.&'#&*.&+'.*2,'8&'$+"4/&M'"<'#&,'$*+,3'(*%,&IO6"4,3'75*%'42&' %.D&']F3' T&' J&#$&' &(' 0+(%8<*4.:' 0' ("4(&,' 8&,' +D/D(%(%"2,' >4%' &26"B*%&2(' #&' G"4I%23' 2",' >4&#>4&,'8"28&+(,'*4'8&+8#&3'&('8&,'.&4@'-&+%*#'-4,%8'U"4+:'

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'''04@' uIG&H,'Z' `<*("23' U*H(*H3' U%13' =4#"' &(' )&/5I:' 0' 8&' e&&H' &2.' .&' #*' #"",&' 9' T*$4D/%&' /+"8<&'.&'6&+B'G*.'(+%/3'&('8&,'+D/;(,'9'#5*++*8<&:'04@',"#",'.&'U*H(*H'&('9'("4,'8&,'8"4/,'.&' /%?:'

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'''0' 1&,' ."8(&4+,3' /"4+' #&4+' *884&%#' &(' #&4+' ?%2&,,&'Z' `<*("23'=4#"3' U%13' T*' -4,,3' T&' ^4%.&3' -%#"4(&3'=&*2OC&G3'U<"1*,3'J*#%3'0%##*%23'04.&3'`<#"D3'`*1%##&3'`#D1&2(3'`*,,*2.+&3'-*+%"2' &('J&4G&4%##&:' ' ' '''0'-4143'd@3'-*+(%8<3'^&B.3'!DI&((&3'-%8<"43'T*I53')%2/%23'-*2"2'&('L#%,&:' ' ' '''0'#*'/+"1"(%"2'`+D/%2'Z'C*2.+%2&3'L#".%&3'C(&/<3'`<#"D3'T*v(%(%*3'`"4,(53'C("4?3'P4%8<3'TD*3' C*2.+*3'P"G%23'=4#%&23',*2,'"4G#%&+'04.+&BM' ' ' '''04@'/"4#"(,3'G%&2'G+%1D,'Z'C*G3'C"/<3'J*+G%&+3'=&??3'S*#3'01%8%&3'V4$#*2.3'_"22&3'-DI*+.3' -*@3'-"+&*43'`<+%,("48<&3'TD2*3'-"+$*2&3')*4#%2&3'J"4+?%"23'm3')%&++&3'V%w%.:' ' '

'''L2?%23' 42&' /&2,D&' D14&' /"4+' ("4,' 8&,' $&2,3' &@(D+%&4+,' 8"11&' 6D("3' >4%' "2(' 8+"%,D' 1"2' 8<&1%2' &2' G"413' $#%,,D' ,4+' #&' 8*++&#*$&' 9' 8*4,&' .4' #%>4%.&' 6*%,,&##&' "4' D(D' (*8#D,' /*+' 1D$*+.&3' G4' .&,' G"%,,"2,' /%D$D&,3' "2(' D(D' GD2%,' /*+' #*' G+",,&' 9' 8<%"((&M' &(' >4%' "2(' 8"2(+%G4D'9'1"2'D/*2"4%,,&1&2('#"+,'.&'8&,'2"1G+&4,&,',"%+D&,M'

(15)
(16)

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" " ,-./0"102",-./0-345"67839025":;<,<89-:;702"0,"-==0402"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"#!" ! /72,0"102"-.90?7-,7<=2"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"#&" ! 7=,9<13@,7<="">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"#(" ! !>" "/0" /-.<9-,<790" 10" :-9-27,</<870" A" 9</0" 0," 7=,090,2" -3" 207=" 1B3=" @-.7=0," ?0,097=-790"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"#*" " """""""!>!>" C32,767@-,7<="10"/-"@90-,7<="1B3="/-.<9-,<790"10":-9-27,</<870"">>>>"#*" """"""""""""""!>!>!>""3D"EFGH"EIGJ"FK"LMNODIHPMQ"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"#*" """"""""""""""!>!>#>""0FNJOMHHKRKDP"LG"LIRNMDK"LBNQPMSMPT"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"$+" """""""""""""""""""""!>!>#>!>"""""/KH"KUNRKDH"EIHHMVFKH"KD"QFMKDPWFK"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"$+" """"""""""""""""""""""""""""!>!>#>!>!>"""""""1MNODIHPMQ"LKH"ENJNHMPIHKH"KUPKJDKH"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"$+" """"""""""""""""""""""""""""!>!>#>!>#>"""""""1MNODIHPMQ"LKH"ENJNHMPIHKH"MDPKJDKH"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"$!" """""""""""""""""""""!>!>#>#>"""""/KH"KUNRKDH"X"YNMJK"JTNFMHKJ"ENJ"GD"FNVIJNPIMJK"HETQMNFMHT"">>>>>>>>>>>>>>>>>>"$%" """"""""""""""!>!>$>""9NEEIJPH"MDPTJZP"HQMKDPMYM[GK\QI]P\VTDTYMQK"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"$)" """""""!>#>" @<=2,7,3,7<="1B3="/-.<9-,<790"10":-9-27,</<870"6<=@,7<==0/"">>>>>>>"%+" """"""""""""""!>#>!>""^NPTJMKF"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"%+" """"""""""""""!>#>#>""-RTDNOKRKDP"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"%#" ! #>" "/02":-9-27,<202"04,09=02"102".<?7=2"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"%$" """""""#>!>" :9020=,-,7<="">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"%$" """"""""""""""#>!>!>""7DYKHPNPMID"ENJ"LKH"EIGU"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"%$" """"""""""""""#>!>#>"",KMODKH"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"%&" """"""""""""""#>!>$>""7DYKHPNPMID"ENJ"LKH"PM[GKH"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"%(" """"""""""""""#>!>%>""8NFKH"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"%)" """"""""""""""#>!>&>"",JIRVMQGFIHK"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"&#" """"""""""""""#>!>'>""1TRILTQMK"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"&$" """"""""""""""#>!>(>""^_MNHKH"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"&&" """""""""""""""""""""#>!>(>!>"""""^_MNHK"MDPKJDK"A"`_EILKJRIHK"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"&&" """""""""""""""""""""#>!>(>#>"""""^_MNHKH"QGPNDTKH"">>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>>"&(" """"""""""""""#>!>)>""1KJRNPMPKH"LGKH"X"LKH"EM[]JKH"LK"LMEPWJKH"`TRNPIE`NOKH"">>>>>>>>>>>>>>>>>>>>>>>>>>>>"&)"

(17)

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(18)

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